Alex Mustard / Midnight snapper (Macolor macularis) shelters in a cavern on a coral reef with white soft corals (Scleronephthya sp.) and solderfish. Vavuu Atoll, Maldives. Indian Ocean

Ernie Janes / Bald eagle (Haliaeetus leucocephalus) portrait, captive, occurs in North America.

Shane Gross / Larval flounder (possibly Laeops Kitahara) in the open ocean at night off Anilao, Philippines.

Nick Upton / Captive reared juvenile White stork (Ciconia ciconia) in flight over the Knepp Estate soon after release, Sussex, UK, August 2019.

Nick Upton / Captive reared juvenile White stork (Ciconia ciconia) with a GPS tracker on its back in flight over the Knepp Estate soon after release, Sussex, UK, August 2019.

Marie Read / American avocet (Recurvirostra americana), adult in breeding plumage foraging by sweeping its bill from side to side through shallow water, Orange County, California, USA, April.

Marie Read / Arctic terns (Sterna paradisaea), two in flight over nesting colony in field of buttercups (Ranunculus sp.), Keflavik, Iceland. Digital canvas expansion.

Marie Read / American oystercatcher (Haematopus palliatus) adult feeding chick, Long Island, New York, USA. June.

Matthew Maran / Delacour's langurs (Trachypithecus delacouri) resting, Van Long Nature Reserve,Ninh Binh, Vietnam. Critically endangered

Cyril Ruoso / Ring-tailed Lemur (Lemur catta), Anja Park, Madagascar

Will Burrard-Lucas / Black and white image of African elephant (Loxodonta africana) Tsavo Conservation Area, Kenya. Taken with a remote camera buggy / BeetleCam. Editorial use only.

Will Burrard-Lucas / Black and white image of African elephant (Loxodonta africana) herd with calf at waterhole, seen through adults legs. Tsavo Conservation Area, Kenya. Taken with a remote camera buggy / BeetleCam. Edit...

Will Burrard-Lucas / Black and white image of African elephant (Loxodonta africana) bulls males facing off against each other, Tsavo Conservation Area, Kenya. Editorial use only.

Will Burrard-Lucas / Black and white image of African elephant (Loxodonta africana) mud bathing, Tsavo Conservation Area, Kenya. Editorial use only.

Piotr Naskrecki / Painted Reed Frog (Hyperolius marmoratus) being eaten by Fishing Spider (Pisauridae), Gorongosa National Park, Mozambique

Will Burrard-Lucas / Black and white image of African elephant (Loxodonta africana) herd, females with extremely long tusks, Tsavo Conservation Area, Kenya. Editorial use only.

Will Burrard-Lucas / Black and white image of African elephant (Loxodonta africana) herd at water hole, female with extremely long tusks, Tsavo Conservation Area, Kenya. Editorial use only.

Will Burrard-Lucas / Black and white image of African elephant (Loxodonta africana) matriarch with herd, Tsavo Conservation Area, Kenya. Editorial use only.

Will Burrard-Lucas / Black and white image of African elephant (Loxodonta africana) female with extremely long tusks, Tsavo Conservation Area, Kenya. Editorial use only.

Will Burrard-Lucas / Black and white image of African elephant (Loxodonta africana) female with extremely long tusks, Tsavo Conservation Area, Kenya. Editorial use only.

Will Burrard-Lucas / Black and white image of African elephant (Loxodonta africana) bull with extremely long tusks, Tsavo Conservation Area, Kenya. Editorial use only.

Will Burrard-Lucas / Black and white image of African elephant (Loxodonta africana) herd at waterhole, Tsavo Conservation Area, Kenya. Editorial use only.

Bernard Castelein / Beech woodland (Fagus sylvatica) with sun rays shining through, black and white image. Peerdsbos, Brasschaat, Belgium

Danny Green / Bald eagle (Haliaeetus leucocephalus) in flight, Alaska, USA, February

Danny Green / Bald eagle (Haliaeetus leucocephalus) in flight, Alaska, USA, February

Edwin Giesbers / Giant panda (Ailuropoda melanoleuca) in snow, captive.

David Pattyn / Black and white ruffed lemur (Varecia variegata variegata) hanging from branch, Vakona island, Andasibe area, Madagascar. Captive.

Scott Leslie / Black-and-white Warbler (Mniotilta varia) male calling in spring, Maine

Agustin Esmoris / Black-throated Mango (Anthracothorax nigricollis) hummingbird feeding on flower nectar, Argentina

Ann & Steve Toon / Pied crows (Corvus albus) perched on horns of antelope skull. Zimanga Private Game Reserve, KwaZulu-Natal, South Africa.

SCOTLAND: The Big Picture / Atlantic salmon (Salmo salar) leaping up a small waterfall in order to reach breeding grounds further upstream, Cairngorms National Park, Scotland, UK, October.

Bence Mate / White-throated Mountain Gem (Lampornis castaneoventris), Green-crowned Brilliant (Heliodoxa jacula) group of three with one in flight, Savegre, Costa Rica

Neil Aldridge / Cheetah (Acinonyx jubatus) staring back over its shoulder, black and white. Save Valley Conservancy, Zimbabwe, August.

Marie Read / Eared grebes (Podiceps nigricollis), pair, one adult carrying food (damselfly) for two chicks riding on the other adult's back, Bowdoin National Wildlife Refuge, Montana, USA

Alex Hyde / Yellow-flecked glassfrogs (Sachatamia albomaculata) mating pair, Osa Peninsula, Costa Rica

Alex Hyde / Yellow-flecked glassfrogs (Sachatamia albomaculata) mating pair, Osa Peninsula, Costa Rica

Jim Brandenburg / Forest on lake shore, Judd Lake, Superior National Forest, Minnesota

Bence Mate / Cormorant (Phalacrocorax carbo) and Great egret (Ardea alba) fighting over fish, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary, January.

Bence Mate / Great egret (Ardea alba) fighting with Cormorant (Phalacrocorax carbo) over fish, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary.

Mark Taylor / Black and white Jack-a-poo, Jack Russell cross Poodle, pup, 8 weeks old, jumping up.

Mark Taylor / Black-and-white kitten, Loona, age 11 weeks, arching back.

Mark Taylor / Black and white Border collie looking sideways at black and white kitten.

Mark Taylor / Black-and-white Springer Spaniel puppy, age 6 weeks.

Wim van den Heever / Bare-faced curassow (Crax fasciolata) portrait of female, Pantanal, Brazil.

Markus Varesvuo / Black grouse (Tetrao / Lyrurus tetrix) flock flying in snow, Suomussalmi, Finland, January.

Erlend Haarberg / Mountain hare (Lepus timidus) sitting in snow at night, Norway, April.

Cheryl-Samantha Owen / Shark spotter, in front of black flag at the Kalk Bay post. It provides a warning to ocean goers that visibility is bad and the shark spotters may not see a white shark if it does enter the bay. West...

Colin Monteath/ Hedgehog House / Clouds and mountains reflected in lake, Lake Matheson, Mount Tasman and Mount Cook, Westland National Park, New Zealand

Pal Hermansen / Inuit sled dogs resting, near Ittoqortoormiit, Scoresbysund, Greenland. Second Prize in the Nature category of the Spider Black and White Photography Competition 2015.

Pal Hermansen / Blizzard on the coast of Lofoten in black and white, Norway, March. Winner of the Natural Adversities category of the Asferico Competition 2015.

Bence Mate / Great egrets (Ardea alba) and Grey heron (Ardea cinerea) squabbling over fish, with some in flight. Blurred motion photograph, Lake Csaj, Pusztaszer, Hungary. February.

Alex Mustard / Nudibranch (Flabellina lineata) in aquarium, taken in the field in Gulen, Norway. North East Atlantic Ocean. Collected for scientific research.

Kirkendall-Spring / Oak trees (Quercus) in the fog, Figueroa Mountain, Los Padres National Forest, California, USA.

Kirkendall-Spring / Squaretop Mountain, Bridger Wilderness of the Wind River Range, Bridger National Forest, Wyoming, USA.

Tony Heald / Hanuman langur (Semnopithecus entellus) eating coconut husk, Bandhavgarh National Park, India. March.

Edwin Giesbers / Crested newt (Triturus carnifex) Gelderland, The Netherlands, April. Winner of the Amphibians category of the Wildlife Photographer of the Year Awards (WPOY) Competition 2015. Winner of the Underwater...

Sven Zacek / Eider ducks (Somateria mollissima) males reflected in a small pond, Longyearbyen, Svalbard, Norway. June.

Richard Du Toit / Black wildebeest (Connochaetus gnou) bull silhouetted at dawn, Rietvlei Nature Reserve, Gauteng Province, South Africa.

Espen Bergersen / Razorbills (Alca torda) in flight, with Atlantic puffin (Fratercula arctica) behind, Hornoya, Norway. March

Espen Bergersen / Kittiwakes (Rissa tridactyla) squabbling on ice. Svalbard, Norway. July

Espen Bergersen / Gyrfalcon (Falco rusticolus) and Raven (Corvus corax) in flight, Hornoya bird cliff, Finnmark, Norway. March

Espen Bergersen / Common guillemot (Uria aalge) calling, bridled guillemot morph with white ring around the eye, Hornoya bird cliff, Finnmark, Norway. March.

Fiona Rogers / Dusky leaf monkey (Trachypithecus obscurus) juvenile portrait . Khao Sam Roi Yot National Park, Thailand. March 2015.

Anup Shah / Dusky leaf monkey (Trachypithecus obscurus) juvenile yawning . Khao Sam Roi Yot National Park, Thailand. March 2015.

Danny Green / Japanese cranes (Grus japonensis) displaying in snow, Hokkaido, Japan, February

Danny Green / Japanese crane (Grus japonensis) displaying, Hokkaido, Japan, February

Ole Jorgen Liodden / Magellanic penguin (Spheniscus magellanicus) profile portrait, Falkland Islands, November.

MYN / Marc Pihet / Common wasp (Vespula vulgaris), Maine-et-Loire, France, November. meetyourneighbours.net project

Aflo / Stray cat, with black and white face markings, sitting on bench, Aichi, Japan.

Markus Varesvuo / Black-throated diver (Gavia arctica) on water, Finland, May.

Markus Varesvuo / Black-throated diver (Gavia arctica) on water, Finland, May.

Markus Varesvuo / Black grouse (Lyrurus tetrix) mating in snow, Utajarvi, Finland, May.

Markus Varesvuo / Willow grouse (Lagopus lagopus), Kiilopaa, Inari, Finland, January.

Floris van Breugel / Black and white photograph of Ice crystals up to 12 inches long along a small creek in a remote valley, Brooks Range, Gates of the Arctic National Park, Alaska, August 2014.

Floris van Breugel / Black and white photograph of humpback whale (Megaptera novaeangliae) tail slapping at surface, Glacier Bay National Park, Alaska, USA, August 2014.

Aflo / Deep sea squid (Chiroteuthis imperator) dead specimen floating, Osezaki, Shizuoka, Izu, Japan.

Espen Bergersen / Adult male King eider (Somateria spectabilis) portrait from behind also showing triangular black sails on the back and white sides of rump. Batsfjord, Norway, March.

Pete Oxford / Guereza colobus monkeys (Colobus guereza) in tree, Lango Bai, Republic of Congo (Congo-Brazzaville), Africa.

Bence Mate / Great egret (Ardea alba) and Grey heron (Ardea cinerea) fighting over a fish, Lake Csaj, Kiskunsagi National Park, Hungary

Fiona Rogers / Red-tail Monkey (Cercopithecus ascanius) in tree, Kakamega Forest Reserve, Kenya

Mark Raycroft / Boston Terrier (Canis familiaris) jumping

Mark Raycroft / Border Collie (Canis familiaris) jumping over branch

Markus Varesvuo / Steller's Eagle (Haliaeteus pelagicus) in flight over snow, Hokkaido, Japan, February

Markus Varesvuo / Steller's Eagle (Haliaeteus pelagicus) flying past snowy mountain peak, Hokkaido, Japan, February

Markus Varesvuo / Red-crowned Crane (Grus japonensis) in flight, Hokkaido, Japan, February

Mark Taylor / Red toy poodle looking up at camera an sticking out tongue.

Richard Robinson / William Trubridge diving - a world champion free diver who is campaigning to protect the Critically Endangered Maui's dolphin (Cephalorhynchus hectori maui) endemic to New Zealand. January 2008. Edi...

Markus Varesvuo / Silhouette of Hooded Crow (Corvus cornix) against full moon, Helsinki, Finland, December

Ammonite / Group of African lion (Panthera leo) cubs playing with their mother and another lioness, footage taken at night using starlight camera technology, Masai Mara, Kenya.

Ammonite / Group of African lion (Panthera leo) cubs play fighting with each other, footage taken at night using starlight camera technology, Masai Mara, Kenya.

Ammonite / Group of African lion (Panthera leo) cubs play fighting with each other, footage taken at night using starlight camera technology, Masai Mara, Kenya.

Ammonite / African lion (Panthera leo) cub playing with mother before walking away, , footage taken at night using starlight camera technology, Masai Mara, Kenya.

Ammonite / African lioness (Panthera leo) growling and swatting at a sub-adult male approaching a kill, footage taken at night using thermal camera technology, Masai Mara, Kenya.

Ammonite / Group of sub-adult African lions (Panthera leo) chasing and jumping on a mother Hippopotamus (Hippopotamus amphibius) and her calf, footage taken at night using infrared camera technology, Masai Mara,...

Ammonite / Sub-adult African lioness (Panthera leo) chasing and jumping onto the back of a Hippopotamus (Hippopotamus amphibius), footage taken at night using infrared camera technology, Masai Mara, Kenya.