Alex Mustard / Commensal emperor shrimp (Periclimenes imperator) moves across the colourful red patterned surface of its host Candycane sea cucumber (Thelenota rubralineata). Misool, Raja Ampat, West Papua, Indonesi...
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Alex Mustard / Reef lizardifsh (Synodus variegatus) at a cleaning station, being cleaned by Holthuis anemone shrimps (Ancylomenes holthuisi). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.
John Chan / Crenulated fiddler crab (Uca crenulata) waving its major claw at another crab, part of a courtship display, Southern California, USA, June.
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John Chan / Crenulated fiddler crab (Uca crenulata) feeding on detritus, Southern California, USA, June.
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John Shaw / Sally lightfoot crabs (Grapsus grapsus) on the beach at Puerto Egas, Santiago Island, Galapagos, May.
Andy Rouse / Sally lightfoot crab, (Grapsus grapsus), feeding on marine iguana parasites and dead skin, Galapagos
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Georgette Douwma / Boxer Crab (Lybia tessellata) with sea anemones in its claws for defense. Lembeh Strait, North Sulawesi, Indonesia.
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Nick Hawkins / European green crab (Carcinus maenas), an invasive species in North America, Kejimkujik Seaside National Park, Nova Scotia, Canada. July.
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SCOTLAND: The Big Picture / Close up of a Long clawed squat lobster (Munida rugosa) in Loch Carron, January 2016.