Nick Gordon / Silky marmosets {Callithrix humeralifer chrysoleuca} aggression, Amazonia, Brazil
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Anup Shah / Juvenile Tawny or African Sea Eagle defending flamingo carcass from Marabou stork {Lepitoptilos crumeniferous}
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Richard Du Toit / Burchell's Zebra (Equus burchellii) males fighting, Rietvlei Nature Reserve, Gauteng, South Africa
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Richard Du Toit / Burchell's Zebra (Equus burchellii) males fighting, Rietvlei Nature Reserve, Gauteng, South Africa
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Paul Hobson / Arctic tern (Sterna paradisaea) two fighting in the air, Scotland, UK July
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Denis-Huot / Masai Giraffe (Giraffa camelopardalis tippelskirchi), males fighting. Masai-Mara game reserve, Kenya.
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Igor Shpilenok / Male Caucasian turs (Capra caucasica) rutting with Dykh-tau Mountain in the background, Kabardino-Balkarsky Zapovednik / Reserve, Russia, July
Anup Shah / Mature male gelada (Theropithecus gelada) fighting head-on, upper lip curled back revealing teeth. Simien Mountains National Park, Ethiopia.
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David Pattyn / Red foxes (Vulpes vulpes) two fighting, probably over territory, The Netherlands, August
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Juan Carlos Munoz / Great Egrets (Ardea alba) territorial fight above nest colony, Everglades National Park, Florida, USA, February.
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Juan Carlos Munoz / Great Egrets (Ardea alba) territorial dispute above nest colony, Everglades National Park, Florida, USA, February.
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Martin Dohrn / Maricopa harvester ant (Pogonomyrmex maricopa) fighting a different species of ant (Aphaenogaster cockerelli), over territory Arizona, USA
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Andrew Parkinson / Redwings (Turdus iliacus) squabbling over an apple in snow. Derbyshire, UK, February. British Wildlife Photographer of the Year (BWPA) competition 2012, 'Animal Behaviour' category. (Non-ex) Winner of...
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Andy Rouse / European hares (Lepus europaeus) boxing, UK
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Andy Rouse / European hares (Lepus europaeus) boxing, UK
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Paul Williams / Tourists watching two wild Jaguar cubs fight on riverbank whilst third looks on from above (Panthera onca) from boat on river, Pantanal, Pocone, Brazil. No release available.
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Andy Rouse / 2020VISION / European Hares (Lepus europaeus) boxing, female on right. Wales, UK, June. Sequence 2 of 2. Did you know? A group of hares is known as a drove.
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Andy Rouse / 2020VISION / European Hares (Lepus europaeus) boxing, female on right. Wales, UK, June. Sequence 1 of 2.
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Andy Rouse / Agile wallabies (Macropus agilis) sparring and fighting, Bamarru Plains, Northern Territories, Australia
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Andy Rouse / Agile wallabies (Macropus agilis) sparring and fighting, Bamarru Plains, Northern Territories, Australia
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Charlie Summers / Plains Zebra (Equus quagga) in a dominance fight with dust flying, Lake Magadi, Serengeti, Tanzania.
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Edwin Giesbers / Alpine marmots (Marmota marmota) fighting, Hohe Tauern National Park, Austria, July
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Carol Walker / Wild horses / Mustangs, two stallions play fighting, Pryor Mountains, Montana, USA, July 2010
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Kim Taylor / Two Siskins (Carduelis spinus) fighting, Surrey, UK, March
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Mark Hamblin / 2020VISION / Goldfinches (Carduelis carduelis) squabbling over teasel seeds in winter. Hope Farm RSPB reserve, Cambridgeshire, England, UK, February. Crop of 01390677
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Mark Hamblin / 2020VISION / Mountain Hares (Lepus timidus) boxing in winter. Cairngorms National Park, Scotland, UK, January. Did you know? Although they are normally solitary, Mountain hares can gather in groups of up to 20 dur...
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Danny Green / 2020VISION / Two European river otters (Lutra lutra) fighting over a fish, Isle of Mull, Inner Hebrides, Scotland, UK, December
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Danny Green / 2020VISION / European river otters (Lutra lutra) play fighting amongst seaweed, Isle of Mull, Inner Hebrides, Scotland, UK, December
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Colin Varndell / Greenfinches (Carduelis chloris) squabbling in flight, Dorset, UK, February
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George Sanker / Bison (Bison bison) two males head to head, Yellowstone NP, Wyoming, USA
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Orsolya Haarberg / Two Black grouse (Tetrao tetrix) males displaying at lek, Bergslagen, Sweden, April
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Erlend Haarberg / Two Common redpolls (Carduelis flammea) fighting in flight, Norway, March
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Erlend Haarberg / Two Arctic terns (Sterna paradisaea) in flight fighting, Iceland, August
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Erlend Haarberg / Two Mountain hares (Lepus timidus) fighting, Sor-Trondelag, Norway, April
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Erlend Haarberg / Two Mountain hares (Lepus timidus) fighting, Sor-Trondelag, Norway, April
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Rolf Nussbaumer / Northern mockingbird (Mimus polyglottos) adult fighting with Golden-fronted Woodpecker (Melanerpes aurifrons) female over Texas Prickly Pear Cactus (Opuntia lindheimeri) Dinero, Lake Corpus Christi, S...
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Ben Hall / Red deer (Cervus elaphus) stags fighting at dusk during rutting season, Cheshire, UK October
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Eric Dragesco / Two Przewalski's / Takhi Horses (Equus ferus prezewalski) fighting, Hustai Nuru National Park, Mongolia
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Carol Walker / Mustang / wild horses, second year cremello colt Claro play fighting with bay, McCullough Peak herd, Wyoming, USA, July 2009
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Carol Walker / Mustang / wild horses, two yearling cremello colts Claro and Cremesso playing with a bay, McCullough Peak herd, Wyoming, USA, June 2008
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Marie Read / Two Black-capped chickadees (Poecile atricapillus) in mid-air fight by a sunflower, New York, USA, February
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Gerrit Vyn / Dall sheep (Ovis dalli) rams head butting, one behaviour by which males establish dominance during the Autumn mating season. Denali National Park, Alaska, USA
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Laurie Campbell / Goldfinches (Carduelis carduelis) squabbling near seed feeder in garden, Berwickshire, Scotland, April
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Edwin Giesbers / Two Red fox (Vulpes vulpes) cubs, four months, play fighting, The Netherlands, June
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Laurie Campbell / Goldfinches (Carduelis carduelis) squabbling near seed feeder, Berwickshire, Scotland, April
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Sergey Gorshkov / Ruffs displaying and fighting (Philomachus pugnax) Agapa River, Taimyr Peninsula, Siberia, Russia
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Bertie Gregory / 2020VISION / Red deer (Cervus elaphus) stags fighting during rut, Richmond Park, London, England, UK, October. Did you know? Richmond Park deer herds are descended from historic medieval royal hunting stock.
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Bertie Gregory / 2020VISION / Red deer (Cervus elaphus) stags fighting during rut, with blocks of flats in the background, Richmond Park, London, England, UK, October
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Bertie Gregory / 2020VISION / Red deer (Cervus elaphus) stags fighting during rut, Richmond Park, London, England, UK, October
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Markus Varesvuo / Horned / Slavonian grebe (Podiceps auritus) two individuals fighting in water, Finland May
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Kristel Richard / During the Festa do Colete Encarnado (Red Waistcoat Festival), a bull running festival, a traditionally dressed 'cavalheiro' delivers the 'banderilla' mounted on his Lusitano stallion in the bullring...
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Juan Carlos Munoz / Two White Tailed Sea Eagle (Haliaeetus albicilla) fighting in acrobatic flight as Raven (Corvus corax) looks on. Scandinavia, April. 5th, 2010 Euronatur Wildlife photographer competition.
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Juan Carlos Munoz / Two White Tailed Sea Eagle (Haliaeetus albicilla) fighting in acrobatic flight as Raven (Corvus corax) looks on and a Herring Gull (Larus argentus) flies past. Scandinavia, April.
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Sven Zacek / Two Black Grouse (Tetrao tetrix) fighting at a lek. Virumaa, Estonia, Europe, April.
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Anup Shah / Caracal (Caracal caracal) two six month kittens play fighting, Masai Mara National Reserve, Kenya, August
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Anup Shah / Caracal (Caracal caracal) two six month kittens play fighting, Masai Mara National Reserve, Kenya, August
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Andrew Parkinson / 2020VISION / Great crested grebe (Podiceps cristatus) adult aggressively pursues another in a territorial dispute, Derbyshire, UK, March
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Andy Rouse / Two Common Zebra (Equus quagga) fighting. Masai Mara, Kenya.
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Colin Varndell / Male Greenfinches (Carduelis chloris) squabbling in flight. Dorset, UK, March.
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Terry Whittaker / 2020VISION / Two Red deer (Cervus elaphus) stags fighting, rutting season, Bushy Park, London, UK, October
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Terry Whittaker / 2020VISION / Red deer (Cervus elaphus) stag chasing another during a fight, rutting season, Bushy Park, London, UK, October
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Terry Whittaker / 2020VISION / Two Red deer (Cervus elaphus) stags fighting, rutting season, Bushy Park, London, UK, October (This image may be licensed either as rights managed or royalty free.)
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Terry Whittaker / 2020VISION / Two Fallow deer (Dama dama) bucks during fight, rutting season, Richmond Park, London, UK, October (This image may be licensed either as rights managed or royalty free.)
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Ole Jorgen Liodden / Beisa oryx (Oryx beisa) fighting in rut. Samburu, Kenya, December.
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Ole Jorgen Liodden / Galapagos Ghost Crab (Ocypode gaudichaudii) fighting on the beach. Galapagos
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Andrew Parkinson / 2020VISION / Marsh harrier (Circus aeruginosus) three sub-adult siblings squabbling in flight in the last rays of evening sunlight, Norfolk, UK, August
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Terry Whittaker / 2020VISION / Konik horse (Equus caballus) two stallions fighting, Wicken Fen, Cambridgeshire, UK, June
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Terry Whittaker / 2020VISION / Konik horse (Equus caballus) two stallions fighting, Wicken Fen, Cambridgeshire, UK, June
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Dave Watts / Eastern grey kangaroo (Macropus giganteus) two males boxing, Australian Capital Territory, Australia, December
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Mary McDonald / Common Zebra (Equus quagga) stallions fighting. Naabi Plains, Serengeti National Park, Tanzania.
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Mary McDonald / Common Zebra (Equus quagga) stallions fighting. Naabi Plains, Serengeti National Park, Tanzania.
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Loic Poidevin / Ruff (Philomachus pugnax) two rival males displaying, fighting, Varanger, Norway, June
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Loic Poidevin / Ruff (Philomachus pugnax) two rival males displaying, fighting, Varanger, Norway, June
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Ernie Janes / Two male Mute swans (Cygnus olor) displaying on water in territorial dispute, UK
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Mark Hamblin / 2020VISION / Black grouse (Tetrao tetrix) males fighting at lek site, Cairngorms National Park, Grampian, Scotland, UK, April
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Mark Hamblin / 2020VISION / Black grouse (Tetrao tetrix) males fighting at lek site, Cairngorms National Park, Grampian, Scotland, UK, April
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Andy Rouse / Two Grizzly bears (Ursus arctos horribilis) male (right) and female fighting over salmon, Katmai NP, Alaska, USA, August
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Andy Rouse / Two Grizzly bears (Ursus arctos horribilis) male (right) and female fighting over salmon, Katmai NP, Alaska, USA, August
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Anup Shah / Adolescent male Lions (Panthera leo) play fighting, Masai Mara National Reserve, Kenya, March
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Bence Mate / Confrontation between a great egret (Ardea alba) and grey heron (Ardea cinerea), Lake Csaj, Kiskunsagi National Park, Hungary.
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Paul Hobson / Brown bear (Ursus arctos) two fighting, Finland, June
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Daniel Heuclin / Cocks fighting in cock fight, Guimaras island, Philippines, February 2007
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Terry Whittaker / 2020VISION / Two Grey herons (Ardea cinerea) squabbling over fish fed to them by visitors,River Tame, Reddish Vale Country Park, Stockport, Greater Manchester, UK, December 2010
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Terry Whittaker / 2020VISION / Three Grey herons (Ardea cinerea) fighting in Regent's Park, London, UK, April 2011.
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Fabrice Cahez / Northern / Hen Harrier (Circus cyaneus) and Kestrel (Falco tinnunculus) below, fighting in flight. Vosges, France, February.
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Nature Production / Two male Babirusa fighting (Babyrousa babyrussa) Sulawesi, Indonesia, vulnerable species
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Markus Varesvuo / Black Grouse (Tetrao tetrix) fighting at a lek. Liminka, Finland, April. Magic Moments book plate, page 45.
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Markus Varesvuo / White-tailed Sea Eagle (Haliaeetus albicilla) being attacked by a Common Gull (Larus canus) in flight. Norway, July. Magic Moments book plate, page 50.