Staffan Widstrand / Elephant dung beetles (Dynastinae) St Lucia wetlands National Park, South Africa

Hiroya Minakuchi / Rhinoceros Beetle (Xylotrupes gideon), Malaysia

Piotr Naskrecki / Rhinoceros Beetle (Dynastinae) male, Tapanti National Park, Cartago, Costa Rica

Piotr Naskrecki / Rhinoceros Beetle (Dynastinae), La Selva Biological Research Station, Heredia, Costa Rica

Stephen Dalton / Close up of claw of Rhinoceros beetle (Dynastinae)

Murray Cooper / Rhinoceros Beetle (Enema pan) showing horns, Ecuador

Murray Cooper / Rhinoceros beetle (Enema pan) on rainforest floor, Ecuador

James Christensen / Rhinoceros Beetle (Megasoma sp), Andes, Ecuador

Tom Vezo / Rhinoceros beetle, Tortugero NP, Costa Rica

Tom Vezo / Rhinoceros beetle, Tortugero NP, Costa Rica

Doug Wechsler / Elephant beetle (Dynastinae) male Philippines Mount Apo NP

Tom Vezo / Rhinoceros Beetle (Dynastinae) amid tree leaves, Tortuguero National Park, Costa Rica

Claus Meyer / Rhinoceros Beetle (Megasoma actaeon) male held in human hand, Amazon ecosystem, Brazil

Mitsuhiko Imamori / Rhinoceros Beetle (Xylotrupes gideon) pair fighting, Asia

Mitsuhiko Imamori / Rhinoceros Beetle (Xylotrupes gideon) pair fighting, Asia