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Freshwater pearl mussels (Margaritifera margaritifera) on river bed, Ennerdale Valley, Lake District NP, Cumbria, England, UK, October 2011
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Linda Pitkin / 2020VISION / Freshwater pearl mussels (Margaritifera margaritifera) on river bed, Ennerdale Valley, Lake District NP, Cumbria, England, UK, October 2011
Bitterling {Rhodeus sericeus} laying eggs into mussel, Europe
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Neil Bromhall / Bitterling {Rhodeus sericeus} laying eggs into mussel, Europe
Freshwater pearl mussels (Margaritifera margaritifera) on river bed, Ennerdale Valley, Lake District NP, Cumbria, England, UK, October 2011
01370131
Linda Pitkin / 2020VISION / Freshwater pearl mussels (Margaritifera margaritifera) on river bed, Ennerdale Valley, Lake District NP, Cumbria, England, UK, October 2011
Freshwater pearl mussels (Margaritifera margaritifera) on river bed, Ennerdale Valley, Lake District NP, Cumbria, England, UK, October 2011
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Linda Pitkin / 2020VISION / Freshwater pearl mussels (Margaritifera margaritifera) on river bed, Ennerdale Valley, Lake District NP, Cumbria, England, UK, October 2011
Freshwater pearl mussels (Margaritifera margaritifera) on river bed, Ennerdale Valley, Lake District NP, Cumbria, England, UK, October 2011. 2020VISION Book Plate.
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Linda Pitkin / 2020VISION / Freshwater pearl mussels (Margaritifera margaritifera) on river bed, Ennerdale Valley, Lake District NP, Cumbria, England, UK, October 2011. 2020VISION Book Plate.
Freshwater pearl mussels (Margaritifera margaritifera) on river bed, Ennerdale Valley, Lake District NP, Cumbria, England, UK, October 2011
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Linda Pitkin / 2020VISION / Freshwater pearl mussels (Margaritifera margaritifera) on river bed, Ennerdale Valley, Lake District NP, Cumbria, England, UK, October 2011
Freshwater pearl mussels (Margaritifera margaritifera) on river bed, Ennerdale Valley, Lake District NP, Cumbria, England, UK, October 2011
01370134
Linda Pitkin / 2020VISION / Freshwater pearl mussels (Margaritifera margaritifera) on river bed, Ennerdale Valley, Lake District NP, Cumbria, England, UK, October 2011
Freshwater pearl mussel (Margaritifera margaritifera) on river bed, Ennerdale Valley, Lake District NP, Cumbria, England, UK, October 2011
01370129
Linda Pitkin / 2020VISION / Freshwater pearl mussel (Margaritifera margaritifera) on river bed, Ennerdale Valley, Lake District NP, Cumbria, England, UK, October 2011
Freshwater pearl mussel (Margaritifera margaritifera) on river bed, Ennerdale Valley, Lake District NP, Cumbria, England, UK, October 2011
01370128
Linda Pitkin / 2020VISION / Freshwater pearl mussel (Margaritifera margaritifera) on river bed, Ennerdale Valley, Lake District NP, Cumbria, England, UK, October 2011
Freshwater pearl mussels (Margaritifera margaritifera) on river bed, Ennerdale Valley, Lake District NP, Cumbria, England, UK, October 2011
01370126
Linda Pitkin / 2020VISION / Freshwater pearl mussels (Margaritifera margaritifera) on river bed, Ennerdale Valley, Lake District NP, Cumbria, England, UK, October 2011
Endemic Mussels (Dreissena presbensis / stankovici) on lake bed at -36 meters, Albania, Eastern Europe, May
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Willem Kolvoort / Endemic Mussels (Dreissena presbensis / stankovici) on lake bed at -36 meters, Albania, Eastern Europe, May
Underwater rock with endemic Mussels (Dreissena presbensis / stankovici) and Sponges (Spongilla stankovici) Albania, Eastern Europe, May
01355746
Willem Kolvoort / Underwater rock with endemic Mussels (Dreissena presbensis / stankovici) and Sponges (Spongilla stankovici) Albania, Eastern Europe, May
Lake bed with endemic Sponge (Porifera), Shrimps (Ohridaspongia rotunda) and Mussels, Albania, Eastern Europe, May
01355745
Willem Kolvoort / Lake bed with endemic Sponge (Porifera), Shrimps (Ohridaspongia rotunda) and Mussels, Albania, Eastern Europe, May
endemic Mussels (Dreissena presbensis / stankovici), Albania, Eastern Europe, May
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Willem Kolvoort / endemic Mussels (Dreissena presbensis / stankovici), Albania, Eastern Europe, May
endemic Mussels (Dreissena presbensis / stankovici), Albania, Eastern Europe, May
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Willem Kolvoort / endemic Mussels (Dreissena presbensis / stankovici), Albania, Eastern Europe, May
Endemic Mussel (Dreissena presbensis / stankovici), Albania, Eastern Europe, May
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Willem Kolvoort / Endemic Mussel (Dreissena presbensis / stankovici), Albania, Eastern Europe, May
Underwater rock with endemic Mussels (Dreissena presbensis / stankovici) and Sponges (Spongilla stankovici) Albania, Eastern Europe, May
01355738
Willem Kolvoort / Underwater rock with endemic Mussels (Dreissena presbensis / stankovici) and Sponges (Spongilla stankovici) Albania, Eastern Europe, May
Underwater rock with endemic Mussels (Dreissena presbensis / stankovici) and Sponges (Spongilla stankovici) Albania, Eastern Europe, May
01355737
Willem Kolvoort / Underwater rock with endemic Mussels (Dreissena presbensis / stankovici) and Sponges (Spongilla stankovici) Albania, Eastern Europe, May
Colony of Zebra mussels (Dreissena polymorpha) in flooded quarry, a non-native, invasive species, Stoney Cove, Stoney Stratford, Leicestershire, UK, December
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Alex Mustard / 2020VISION / Colony of Zebra mussels (Dreissena polymorpha) in flooded quarry, a non-native, invasive species, Stoney Cove, Stoney Stratford, Leicestershire, UK, December
Freshwater / river mussel (Unio tumidus) shells on the bank of a small rhyne / drainage ditch in the Han Wall RSPB National Nature Reserve, England, December 2009.
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Toby Roxburgh / Freshwater / river mussel (Unio tumidus) shells on the bank of a small rhyne / drainage ditch in the Han Wall RSPB National Nature Reserve, England, December 2009.
Dwarf Wedge Mussel (left), Eastern Elliptio (top), and Eastern Lampmussel (right.) found in the Ashuelot River. Keene, New Hampshire, USA, September 2006.
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Jerry Monkman / Dwarf Wedge Mussel (left), Eastern Elliptio (top), and Eastern Lampmussel (right.) found in the Ashuelot River. Keene, New Hampshire, USA, September 2006.
Mussel researcher Ethan Nedeau holding three species of freshwater Mussels (Mytillus genus) from the Ashuelot River. Keene, New Hampshire, USA, September 2006.
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Jerry Monkman / Mussel researcher Ethan Nedeau holding three species of freshwater Mussels (Mytillus genus) from the Ashuelot River. Keene, New Hampshire, USA, September 2006.
Freshwater pearl mussel {Margaritifera margaritifera} in Ballinderry River, County Tyron, Republic of Ireland, March
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Robert Thompson / Freshwater pearl mussel {Margaritifera margaritifera} in Ballinderry River, County Tyron, Republic of Ireland, March
Freshwater pearl mussel {Margaritifera margaritifera} in Ballinderry River, County Tyron, Republic of Ireland, March
01271886
Robert Thompson / Freshwater pearl mussel {Margaritifera margaritifera} in Ballinderry River, County Tyron, Republic of Ireland, March
Zebra Mussels {Dreissena polymorpha}  Lower Lough Erne, County Fermanagh, Northern Ireland, UK
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Robert Thompson / Zebra Mussels {Dreissena polymorpha} Lower Lough Erne, County Fermanagh, Northern Ireland, UK
Zebra Mussels {Dreissena polymorpha} Lower Lough Erne, County Fermanagh, Northern Ireland, UK
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Robert Thompson / Zebra Mussels {Dreissena polymorpha} Lower Lough Erne, County Fermanagh, Northern Ireland, UK
Freshwater pearl mussel (Margaritifera margaritifera) on river bed, Umeälven tributary, Sweden, July 2008
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Wild Wonders of Europe / Roggo / Freshwater pearl mussel (Margaritifera margaritifera) on river bed, Umeälven tributary, Sweden, July 2008
Bitterling {Rhodeus sericeus} laying eggs into mussel, Europe
01077079
Neil Bromhall / Bitterling {Rhodeus sericeus} laying eggs into mussel, Europe
Oily shiner {Sarcocheilichthys variegatus} inserting its ovipositor and laying eggs in Asian Freshwater Mussel {Anodonta (Sinanodonta) woodiana} captive, Japan
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Nature Production / Oily shiner {Sarcocheilichthys variegatus} inserting its ovipositor and laying eggs in Asian Freshwater Mussel {Anodonta (Sinanodonta) woodiana} captive, Japan
Freshwater pearl mussels (Margaritifera margaritifera) in river, East Latvia
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Willem Kolvoort / Freshwater pearl mussels (Margaritifera margaritifera) in river, East Latvia
Underwater view of Greater european pea clam(Pisidium amnicum) in garden pond, Holland
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Willem Kolvoort / Underwater view of Greater european pea clam(Pisidium amnicum) in garden pond, Holland
Swan mussel (Anodonta cygnaea) walking along river bed floor, IJsselmeer, Holland
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Willem Kolvoort / Swan mussel (Anodonta cygnaea) walking along river bed floor, IJsselmeer, Holland
Freshwater pearl mussel (Margaritifera margaritifera) inlet opening, East Latvia
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Willem Kolvoort / Freshwater pearl mussel (Margaritifera margaritifera) inlet opening, East Latvia
Entrance and discharge openings of the Swan mussel (Anodonta cygnaea) sand winning pit, Holland
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Willem Kolvoort / Entrance and discharge openings of the Swan mussel (Anodonta cygnaea) sand winning pit, Holland
Semi-aquatic weevil (Pachyrhinus / Bagous elegans) on Swan mussel (Anodon cygnaea), Lake Naarden, Holland
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Willem Kolvoort / Semi-aquatic weevil (Pachyrhinus / Bagous elegans) on Swan mussel (Anodon cygnaea), Lake Naarden, Holland
Freshwater pearl mussels (Margaritifera margaritifera) in fast flowing river, East Latvia
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Willem Kolvoort / Freshwater pearl mussels (Margaritifera margaritifera) in fast flowing river, East Latvia
Moss animals / Bryozoans {Cristatella mucedo} (above) and {Plumatella repens} Sand-winning Pit, Holland
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Willem Kolvoort / Moss animals / Bryozoans {Cristatella mucedo} (above) and {Plumatella repens} Sand-winning Pit, Holland
Moss animals (Cristatella mucedo), Zebra mussel (Dreissena polymorpha) and Ramshorn snail (Planorbidae), all on submerged branch, peat bog lake, Holland
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Willem Kolvoort / Moss animals (Cristatella mucedo), Zebra mussel (Dreissena polymorpha) and Ramshorn snail (Planorbidae), all on submerged branch, peat bog lake, Holland
Swan mussels (Anodonta cygnaea) and Perfoliate pondweed (Potamogeton perfoliatus) Czarna Hanca river, Poland
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Willem Kolvoort / Swan mussels (Anodonta cygnaea) and Perfoliate pondweed (Potamogeton perfoliatus) Czarna Hanca river, Poland
Underwater mass of Zebra mussels (Dreissena polymorpha) in sand winning pit, Holland
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Willem Kolvoort / Underwater mass of Zebra mussels (Dreissena polymorpha) in sand winning pit, Holland
A big Zebra mussel (Dreissena polymorpha) with a lot of young mussels around it, maybe offspring, sand winning pit, Holland
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Willem Kolvoort / A big Zebra mussel (Dreissena polymorpha) with a lot of young mussels around it, maybe offspring, sand winning pit, Holland
Close up of Swan mussel (Anodonta cygnaea) the opening with the tentacles is the inlet of the mussel,  sand winning pit, Holland
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Willem Kolvoort / Close up of Swan mussel (Anodonta cygnaea) the opening with the tentacles is the inlet of the mussel, sand winning pit, Holland
Swan mussel (Anodonta cygnaea) on bottom of sand winning pit, Holland
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Willem Kolvoort / Swan mussel (Anodonta cygnaea) on bottom of sand winning pit, Holland
Swan mussel (Anodonta cygnaea) sand-winning pit, Holland
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Willem Kolvoort / Swan mussel (Anodonta cygnaea) sand-winning pit, Holland
Swan mussel (Anodonta cygnaea) creeps through the upper layer of the sandy bottom to look for a better position, Ijsselmeer (former Zuiderzee), Holland
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Willem Kolvoort / Swan mussel (Anodonta cygnaea) creeps through the upper layer of the sandy bottom to look for a better position, Ijsselmeer (former Zuiderzee), Holland
Underwater branches covered in Zebra mussels (Dreissena polymorpha) sand winning pit, Holland
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Willem Kolvoort / Underwater branches covered in Zebra mussels (Dreissena polymorpha) sand winning pit, Holland
Swan mussel (Anodonta cygnaea) with Zebra mussel (Dreissena polymorpha) attached to left side, sand winning pit, Holland
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Willem Kolvoort / Swan mussel (Anodonta cygnaea) with Zebra mussel (Dreissena polymorpha) attached to left side, sand winning pit, Holland
Freshwater pearl mussels (Margaritifera margaritifera) in fast flowing river, East Latvia
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Willem Kolvoort / Freshwater pearl mussels (Margaritifera margaritifera) in fast flowing river, East Latvia
European fingernail clam / Freshwater cockle {Sphaerium corneum} showing siphons, Sussex, UK
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Simon Colmer / European fingernail clam / Freshwater cockle {Sphaerium corneum} showing siphons, Sussex, UK
Swan Mussel (Anodonta cygnaea) with semi-aquatic Weevil. Lake Naarden, Holland. Mussel is filtering water to feed on plankton.
01155858
Willem Kolvoort / Swan Mussel (Anodonta cygnaea) with semi-aquatic Weevil. Lake Naarden, Holland. Mussel is filtering water to feed on plankton.
Zebra mussels (Dreissena polymorpha) feeding in a canal, Zwolle, Holland
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Willem Kolvoort / Zebra mussels (Dreissena polymorpha) feeding in a canal, Zwolle, Holland
Swan Mussel (Anodonta cygnaea) bog at Lake Naarden, Holland. Mussel is filtering water to feed on plankton.
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Willem Kolvoort / Swan Mussel (Anodonta cygnaea) bog at Lake Naarden, Holland. Mussel is filtering water to feed on plankton.
Swan mussel {Anodonta cygnaea} Close up of mantle showing tentacles that sift out debris during feeding, Holland
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Willem Kolvoort / Swan mussel {Anodonta cygnaea} Close up of mantle showing tentacles that sift out debris during feeding, Holland
Greater european pea clams / Pea mussels (Pisidium amnicum) filter feeding in garden pond, Holland
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Willem Kolvoort / Greater european pea clams / Pea mussels (Pisidium amnicum) filter feeding in garden pond, Holland
Zebra mussels {Dreissena polymorpha} attached to aquatic plants, Lough Keye, Republic of Ireland
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Willem Kolvoort / Zebra mussels {Dreissena polymorpha} attached to aquatic plants, Lough Keye, Republic of Ireland
Greater european pea clams / Pea mussels (Pisidium amnicum) filter feeding in garden pond, Holland
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Willem Kolvoort / Greater european pea clams / Pea mussels (Pisidium amnicum) filter feeding in garden pond, Holland
Zebra mussels {Dreissena polymorpha} one adult and several young filter feeding, Holland.
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Willem Kolvoort / Zebra mussels {Dreissena polymorpha} one adult and several young filter feeding, Holland.
Zebra mussels {Dreissena polymorpha} filter feeding, Holland
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Willem Kolvoort / Zebra mussels {Dreissena polymorpha} filter feeding, Holland
Zebra mussels {Dreissena polymorpha} filter feeding, Holland
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Willem Kolvoort / Zebra mussels {Dreissena polymorpha} filter feeding, Holland
Zebra mussels {Dreissena polymorpha} attached to an old steel construction in a new brackish-fresh water lake. Holland
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Willem Kolvoort / Zebra mussels {Dreissena polymorpha} attached to an old steel construction in a new brackish-fresh water lake. Holland
Zebra mussels {Dreissena polymorpha} filter feeding with tentacles preventing discharge from entering. Peat-bog lake, Holland
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Willem Kolvoort / Zebra mussels {Dreissena polymorpha} filter feeding with tentacles preventing discharge from entering. Peat-bog lake, Holland
Swan mussel (Anodonta cygnea) filter feeding on plankton with tentacles to prevent debris from entering, Holland
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Willem Kolvoort / Swan mussel (Anodonta cygnea) filter feeding on plankton with tentacles to prevent debris from entering, Holland
Zebra mussels {Dreissana polymorpha} attached to stone on lake bed. Lough Keye, Republic of Ireland
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Willem Kolvoort / Zebra mussels {Dreissana polymorpha} attached to stone on lake bed. Lough Keye, Republic of Ireland
Close up of Zebra mussel {Dreissena polymorpha} filter feeding with tentacles preventing discharge from entering. Peat-bog lake, Holland
01155955
Willem Kolvoort / Close up of Zebra mussel {Dreissena polymorpha} filter feeding with tentacles preventing discharge from entering. Peat-bog lake, Holland
Swan mussel (Anodonta cygnea) male releasing sperm which will be dispersed by water currents to a female, Holland
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Willem Kolvoort / Swan mussel (Anodonta cygnea) male releasing sperm which will be dispersed by water currents to a female, Holland
Close up of Zebra mussel {Dreissena polymorpha} filter feeding clearly showing the inlet (left) and outlet (right) openings, with tentacles at the inlet opening to prevent rubbish entering, Deep peat-...
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Willem Kolvoort / Close up of Zebra mussel {Dreissena polymorpha} filter feeding clearly showing the inlet (left) and outlet (right) openings, with tentacles at the inlet opening to prevent rubbish entering, Deep peat-...
Zebra mussels {Dreissena polymorpha} attached to a stone on lake floor, Lough Keye, Republic of Ireland
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Willem Kolvoort / Zebra mussels {Dreissena polymorpha} attached to a stone on lake floor, Lough Keye, Republic of Ireland
Swan mussel (Anodonta cygnea) filter feeding on plankton with tentacles to prevent debris from entering, Holland
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Willem Kolvoort / Swan mussel (Anodonta cygnea) filter feeding on plankton with tentacles to prevent debris from entering, Holland
Close up of Zebra mussel {Dreissena polymorpha} filter feeding clearly showing the inlet (lower) and outlet (upper) openings, with tentacles at the inlet opening to prevent rubbish entering, Deep peat...
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Willem Kolvoort / Close up of Zebra mussel {Dreissena polymorpha} filter feeding clearly showing the inlet (lower) and outlet (upper) openings, with tentacles at the inlet opening to prevent rubbish entering, Deep peat...
Zebra mussels {Dreissena polymorpha} attached to a piece of wood on lake floor, Lough Keye, Republic of Ireland
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Willem Kolvoort / Zebra mussels {Dreissena polymorpha} attached to a piece of wood on lake floor, Lough Keye, Republic of Ireland
Freshwater pearl mussel shell (Margaritifera margaritifera) Natural History Museum, Riga, Latvia
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Willem Kolvoort / Freshwater pearl mussel shell (Margaritifera margaritifera) Natural History Museum, Riga, Latvia
Pearls from Freshwater pearl mussel (Margaritifera margaritifera) Natural History Museum, Riga, Latvia
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Willem Kolvoort / Pearls from Freshwater pearl mussel (Margaritifera margaritifera) Natural History Museum, Riga, Latvia
Freshwater pearl mussels (Margaritifera margaritifera) These heavily threatened mussels live in a small river in the forests of Latvia. In most european countries they are extinct.
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Willem Kolvoort / Freshwater pearl mussels (Margaritifera margaritifera) These heavily threatened mussels live in a small river in the forests of Latvia. In most european countries they are extinct.
Child looking at Freshwater Pearl Mussels (Margaritifera margaritifera) Sweden
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Staffan Widstrand / Child looking at Freshwater Pearl Mussels (Margaritifera margaritifera) Sweden
Freshwater Pearl Mussels (Margaritifera margaritifera) Halsingland, Sweden.
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Staffan Widstrand / Freshwater Pearl Mussels (Margaritifera margaritifera) Halsingland, Sweden.
Female Bitterling ovipositing into Mussel while male waits to deposit sperm, sequence 6/6
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Kim Taylor / Female Bitterling ovipositing into Mussel while male waits to deposit sperm, sequence 6/6
Female Bitterling inserts ovipositor into siphon of swan mussel {Rhodeus ocellatus}, sequence 5/6
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Kim Taylor / Female Bitterling inserts ovipositor into siphon of swan mussel {Rhodeus ocellatus}, sequence 5/6
Male Bitterling guides gravid female to Mussel for ovipositing {Rhodeus ocellauts} trailing false ovipositor.
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Kim Taylor / Male Bitterling guides gravid female to Mussel for ovipositing {Rhodeus ocellauts} trailing false ovipositor.
Female Bitterling hangs vertically before ovipositing in Swan mussel {Rhodeus ocellatus}
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Kim Taylor / Female Bitterling hangs vertically before ovipositing in Swan mussel {Rhodeus ocellatus}
Gravid female Bitterling and male inspect freshwater Mussel for spawning, seq 1/6 trailing false ovipositor.
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Kim Taylor / Gravid female Bitterling and male inspect freshwater Mussel for spawning, seq 1/6 trailing false ovipositor.
Male Bitterling guides gravid female to Mussel for ovipositing, sequence 2/6
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Kim Taylor / Male Bitterling guides gravid female to Mussel for ovipositing, sequence 2/6
Female Bitterling hangs vertically before ovipositing in Swan mussel, sequence 3/6 male in attendance trailing false ovipositor.
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Kim Taylor / Female Bitterling hangs vertically before ovipositing in Swan mussel, sequence 3/6 male in attendance trailing false ovipositor.
Female Bitterling dashs down to insert ovipositor into swan mussel, sequence 4/6 male in attendance trailing false ovipositor.
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Kim Taylor / Female Bitterling dashs down to insert ovipositor into swan mussel, sequence 4/6 male in attendance trailing false ovipositor.
Bitterling {Rhodeus sericeus} laying eggs into mussel Europe.
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Neil Bromhall / Bitterling {Rhodeus sericeus} laying eggs into mussel Europe.
Bitterling {Rhodeus sericeus} about to lay eggs into mussel Europe.
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Neil Bromhall / Bitterling {Rhodeus sericeus} about to lay eggs into mussel Europe.
Swan mussel with foot extended {Anodonta cygnaea} Yorkshire, UK
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Paul Hobson / Swan mussel with foot extended {Anodonta cygnaea} Yorkshire, UK
Bitterling {Rhodeus sericeus} about to lay eggs into mussel Europe
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Neil Bromhall / Bitterling {Rhodeus sericeus} about to lay eggs into mussel Europe
Bitterling showing ovipositor tube; lays eggs in mussel. Germany
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Dietmar Nill / Bitterling showing ovipositor tube; lays eggs in mussel. Germany
Freshwater mussel {Inversidens brandti} Shiga, Japan
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Nature Production / Freshwater mussel {Inversidens brandti} Shiga, Japan
Freshwater mussel {Inversiunio reiniana yanagawaensis} Shiga, Japan
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Nature Production / Freshwater mussel {Inversiunio reiniana yanagawaensis} Shiga, Japan
Freshwater mussel {Inversidens japanensis} Shiga, Japan
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Nature Production / Freshwater mussel {Inversidens japanensis} Shiga, Japan
Freshwater mussel {Pseudodon omiensis} Japan
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Nature Production / Freshwater mussel {Pseudodon omiensis} Japan
Bitterling {Acheilognathus rhombeus} pair preparing to spawn in freshwater mussel {Unio douglasiae} Japan
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Nature Production / Bitterling {Acheilognathus rhombeus} pair preparing to spawn in freshwater mussel {Unio douglasiae} Japan
Asian Freshwater Mussel {Anodonta woodiana} Japan
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Nature Production / Asian Freshwater Mussel {Anodonta woodiana} Japan
Freshwater mussel {Unio douglasiae} Japan
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Nature Production / Freshwater mussel {Unio douglasiae} Japan
Freshwater mussel {Lanceolaria grayana cuspidata} Hyogo, Japan
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Nature Production / Freshwater mussel {Lanceolaria grayana cuspidata} Hyogo, Japan