Chris Mattison / Common dor beetle (Geotrupes stercorarius) on moss. Whitelye, Monmouthshire, Wales, UK. March.

Chris Mattison / Common dor beetle (Geotrupes stercorarius) on white background. Whitelye, Monmouthshire, Wales, UK. March.

Chris Mattison / Common dor beetle (Geotrupes stercorarius) on white background. Whitelye, Monmouthshire, Wales, UK. March.

Piotr Naskrecki / Dor Beetle (Geotrupes vernalis), Poland

Kim Taylor / Minotaur Beetle (Typhaeus typhoeus) male. Surrey, England, Controlled conditions

Kim Taylor / Minotaur beetle (Typhaeus typhoeus) male. Surrey, England, UK. October. Controlled conditions

Andy Sands / Dor Beetle (Geotrupes stercorarius) smaller male riding on back of female after mating, Bedfordshire, England, Uk, April

Andy Sands / Dor beetle (Geotrupes stercorarius) mating pair, Bedfordshire, England, Uk, April

Nick Upton / Woodland dor beetle (Anoplotrupes stercorosus), Knapdale, Scotland, UK, May.

Nick Upton / Woodland dor beetle (Anoplotrupes stercorosus) on moss, Knapdale, Scotland, UK, May.

Thomas Hinsche/ BIA / Stag Beetle (Lucanus cervus), Saxony-Anhalt, Germany

MYN / Marc Pihet / Forest dung beetle (Anoplotrupes stercorosus), Maine-et-Loire, France, October, meetyourneighbours.net project

MYN / Marc Pihet / Forest dung beetle (Anoplotrupes stercorosus), Maine-et-Loire, France, October, meetyourneighbours.net project

MYN / Marc Pihet / Forest dung beetle (Anoplotrupes stercorosus), Maine-et-Loire, France, October, meetyourneighbours.net project

MYN / Marc Pihet / Forest dung beetle (Anoplotrupes stercorosus), Maine-et-Loire, France, October, meetyourneighbours.net project

Will Watson / Dor beetle (Geotrupes stercoraurius), walking through bracken and leaf litter, Mortimer Forest, Shropshire, England, UK, October.

Pascal Pittorino / Dor Beetle (Geotrupes stercorarius) on moss, Corsica Island, France, September

John Abbott / Opaque Earth Boring Beetle (Geotrupes opacus) Bastrop County, Texas, USA.

MYN / Emanuele Biggi / Dung beetle (Geotrupes sp.) Italy, September. Meetyourneighbours.net project

MYN / Marko Masterl / Forest Dung Beetle (Anoplotrupes stercorosus) with wing out of elytra, Slovenia, Europe, June Meetyourneighbours.net project

MYN / Marko Masterl / Forest Dung Beetle (Anoplotrupes stercorosus), Slovenia, Europe, June Meetyourneighbours.net project

MYN / Marko Masterl / Forest Dung Beetle (Anoplotrupes stercorosus), Slovenia, Europe, June Meetyourneighbours.net project

MYN / Marko Masterl / Forest Dung Beetle (Anoplotrupes stercorosus), Slovenia, Europe, June Meetyourneighbours.net project

MYN / Marko Masterl / Forest Dung Beetle (Anoplotrupes stercorosus), Slovenia, Europe, June Meetyourneighbours.net project

Thomas Marent / Stag Beetle (Lucanus cervus) males fighting, Switzerland

Thomas Marent / Stag Beetle (Lucanus cervus) males fighting, Switzerland

Thomas Marent / Stag Beetle (Lucanus cervus) males fighting, Switzerland

Thomas Marent / Stag Beetle (Lucanus cervus) males fighting, Switzerland

Thomas Marent / Stag Beetle (Lucanus cervus) male, Switzerland

Thomas Marent / Stag Beetle (Lucanus cervus) male, Switzerland

Thomas Marent / Stag Beetle (Lucanus cervus) male, Switzerland

MYN / Niall Benvie / Dor beetle / Earth-boring dung beetle (Geotrupidae sp.), Scotland, UK, May meetyourneighbours.net project

Thomas Marent / Stag Beetle (Lucanus cervus) male and female, France

Thomas Marent / Stag Beetle (Lucanus cervus) male, France

Ingo Arndt / Stag Beetle (Lucanus cervus) showing large jaws, Germany

Kim Taylor / Dor beetle {Typhaeus typhoeus} male with rabbit dropping / pellet. UK.

Jan Vermeer / Stag Beetle (Lucanus cervus) two males fighting, Europe

Jan Vermeer / Stag Beetle (Lucanus cervus) male on mossy branch, Europe

Jan Vermeer / Stag Beetle (Lucanus cervus) silhouette of male stag beetle on leaf, Europe

Mitsuhiko Imamori / Earth-boring Dung Beetle (Aegus acuminatus) portrait, Asia

Barry Mansell / Scarab beetle (Peltotrupes profundus) endemic to Florida. Controlled conditions.

Barry Mansell / Scarab beetle (Peltotrupes profundus) endemic to Florida. Controlled conditions.

Thomas Marent / Stag Beetle (Lucanus cervus) males fighting, Switzerland

Thomas Marent / Stag Beetle (Lucanus cervus) males fighting, Switzerland

Thomas Marent / Stag Beetle (Lucanus cervus) males fighting, Switzerland

Thomas Marent / Stag Beetle (Lucanus cervus) males fighting, Switzerland

Thomas Marent / Stag Beetle (Lucanus cervus) male, Switzerland

Thomas Marent / Stag Beetle (Lucanus cervus) male, Switzerland

Ashley Cooper / Dor beetle (Anoplotrupes stercorosus) portrait, near Austwick, Yorkshire Dales, UK. May.

Bruce Thomson / Dor beetle (Blackburnium sloanei) female, portrait, Currawinya National Park, Queensland, Australia.

Bruce Thomson / Dor beetle (Blackburnium sloanei) female, portrait, Currawinya National Park, Queensland, Australia.

Alex Hyde / Phoretic mites (Poecilochirus sp.) hitching a lift on the iridescent undercarriage of a Dor beetle (Geotrupes sp.) travelling with the beetle between the dung piles on which it feeds, Wigwell, Derbysh...

Alex Hyde / False-coloured Scanning Electron Micrograph produced at the University of Derby of a Dor beetle head (Geotrupes sp.) part of the dung beetle army that plays a central role in breaking down animal dung...

Alex Hyde / Dor Beetle (Geotrupes stercorarius) Peak District National Park, Derbyshire, UK. July. Focus tacked image.

Alex Hyde / Dor Beetle (Geotrupes stercorarius) Peak District National Park, Derbyshire, UK. July. Focus stacked image, photographed on a white background in mobile feild studio.

Nigel Cattlin / Dung beetle / Dor beetle / Lousy watchman (Geotrupes stercoreus) on soil and debris, Devon, England, UK, August

Nigel Cattlin / Dung beetle / Dor beetle / Lousy watchman (Geotrupes stercoreus) on soil, Devon, England, UK, August