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Humphead parrotfish (Bolbometopon muricatum) shoal spawning in early morning, part of mass aggregation of thousands of fish. Caroline Islands, Palau.
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Tony Wu / Humphead parrotfish (Bolbometopon muricatum) shoal spawning in early morning, part of mass aggregation of thousands of fish. Caroline Islands, Palau.
Male Bullethead parrotfish (Chlorurus sordidus) displaying in late afternoon. Ras Katy, Sinai, Egypt. Gulf of Aqaba, Red Sea.
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Alex Mustard / Male Bullethead parrotfish (Chlorurus sordidus) displaying in late afternoon. Ras Katy, Sinai, Egypt. Gulf of Aqaba, Red Sea.
Close up of mouth of a Stoplight parrotfish (Sparisoma viride) Bahamas, Caribbean
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Jeff Rotman / Close up of mouth of a Stoplight parrotfish (Sparisoma viride) Bahamas, Caribbean
Bullethead parrotfish (Chlororus sordidus) tail. Red Sea, Egypt
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Jeff Rotman / Bullethead parrotfish (Chlororus sordidus) tail. Red Sea, Egypt
Stoplight parrotfish (Sparisoma viride) eating algae on coral reef. Eleuthera, Bahamas.
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Shane Gross / Stoplight parrotfish (Sparisoma viride) eating algae on coral reef. Eleuthera, Bahamas.
Bumphead parrotfish (Bolbometopon muricatum), Yap, Micronesia.
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David Fleetham / Bumphead parrotfish (Bolbometopon muricatum), Yap, Micronesia.
Stoplight parrotfish (Sparisoma viride), male, biting coral rock (feeding on encrusting algae) Bonaire, Leeward Antilles, Caribbean.
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Linda Pitkin / Stoplight parrotfish (Sparisoma viride), male, biting coral rock (feeding on encrusting algae) Bonaire, Leeward Antilles, Caribbean.
Bullethead parrotfish (Chlorurus sordidus) male, close up of eye, whilst sleeping at night. The Alternatives, Sinai, Egypt. Red Sea
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Alex Mustard / Bullethead parrotfish (Chlorurus sordidus) male, close up of eye, whilst sleeping at night. The Alternatives, Sinai, Egypt. Red Sea
Greenthroat or Singapore parrotfish (Scarus prasiognathos), terminal male grazing on algae covered coral boulder,  Andaman Sea, Thailand. December.
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Georgette Douwma / Greenthroat or Singapore parrotfish (Scarus prasiognathos), terminal male grazing on algae covered coral boulder, Andaman Sea, Thailand. December.
Bumphead parrotfish (Bolbometopon muricatum) gather at dawn on a shallow coral reef. Sipadan Island, Sabah, Borneo, Malaysia. Celebes Sea. Vulnerable species.
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Alex Mustard / Bumphead parrotfish (Bolbometopon muricatum) gather at dawn on a shallow coral reef. Sipadan Island, Sabah, Borneo, Malaysia. Celebes Sea. Vulnerable species.
Stoplight parrotfish (Sparisoma viride) on coral reef,  Hol Chan Marine Reserve, Belize.
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Pete Oxford / Stoplight parrotfish (Sparisoma viride) on coral reef, Hol Chan Marine Reserve, Belize.
Greenthroat or Singapore parrotfish (Scarus prasiognathus), large school of terminal males grazing on algae covered coral boulders. Andaman Sea, Thailand.
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Georgette Douwma / Greenthroat or Singapore parrotfish (Scarus prasiognathus), large school of terminal males grazing on algae covered coral boulders. Andaman Sea, Thailand.
Greenthroat or Singapore parrotfish (Scarus prasiognathus), grazing on a granite boulder, with a small Remora (Echeneis naucrates) attached. Andaman Sea, Thailand.
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Georgette Douwma / Greenthroat or Singapore parrotfish (Scarus prasiognathus), grazing on a granite boulder, with a small Remora (Echeneis naucrates) attached. Andaman Sea, Thailand.
Portrait of a female Stoplight parrotfish (Sparisoma viride) hovering above a coral reef. Bucaneer Reef, Cayman Brac, Cayman Islands. Caribbean Sea.
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Alex Mustard / Portrait of a female Stoplight parrotfish (Sparisoma viride) hovering above a coral reef. Bucaneer Reef, Cayman Brac, Cayman Islands. Caribbean Sea.
Stoplight parrotfish (Sparisoma viride), super male (will always remain male), terminal phase.  Bonaire, Netherlands Antilles, Caribbean, Atlantic Ocean.
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Georgette Douwma / Stoplight parrotfish (Sparisoma viride), super male (will always remain male), terminal phase. Bonaire, Netherlands Antilles, Caribbean, Atlantic Ocean.
Stoplight parrotfish (Sparisoma viride)  terminal phase.  Bonaire, Netherlands Antilles, Caribbean, Atlantic Ocean.
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Georgette Douwma / Stoplight parrotfish (Sparisoma viride) terminal phase. Bonaire, Netherlands Antilles, Caribbean, Atlantic Ocean.
Stoplight parrotfish (Sparisoma viride) initial phase.  Bonaire, Netherlands Antilles, Caribbean, Atlantic Ocean.
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Georgette Douwma / Stoplight parrotfish (Sparisoma viride) initial phase. Bonaire, Netherlands Antilles, Caribbean, Atlantic Ocean.
Stoplight parrotfish (Sparisoma viride) initial phase.  Bonaire, Netherlands Antilles, Caribbean, Atlantic Ocean.
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Georgette Douwma / Stoplight parrotfish (Sparisoma viride) initial phase. Bonaire, Netherlands Antilles, Caribbean, Atlantic Ocean.
Stoplight parrotfish (Sparisoma viride)  initial phase turning into terminal phase.  Bonaire, Netherlands Antilles, Caribbean, Atlantic Ocean.
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Georgette Douwma / Stoplight parrotfish (Sparisoma viride) initial phase turning into terminal phase. Bonaire, Netherlands Antilles, Caribbean, Atlantic Ocean.
Stoplight parrotfish (Sparisoma viride), terminal phase, super male. Bonaire, Netherlands Antilles, Caribbean, Atlantic Ocean.
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Georgette Douwma / Stoplight parrotfish (Sparisoma viride), terminal phase, super male. Bonaire, Netherlands Antilles, Caribbean, Atlantic Ocean.
Stoplight parrotfish (Sparisoma viride), terminal phase, grazing on coral rock, shadowed by a Trumpetfish (Aulostomus maculatus) next to a Rocky beauty (Holacanthus tricolor)  Bonaire, Netherlands Ant...
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Georgette Douwma / Stoplight parrotfish (Sparisoma viride), terminal phase, grazing on coral rock, shadowed by a Trumpetfish (Aulostomus maculatus) next to a Rocky beauty (Holacanthus tricolor) Bonaire, Netherlands Ant...
Stoplight parrotfish (Sparisoma viride) terminal phase.  Bonaire, Netherlands Antilles, Caribbean, Atlantic Ocean.
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Georgette Douwma / Stoplight parrotfish (Sparisoma viride) terminal phase. Bonaire, Netherlands Antilles, Caribbean, Atlantic Ocean.
Stoplight parrotfish (Sparisoma viride) initial phase.  Bonaire, Netherlands Antilles, Caribbean, Atlantic Ocean.
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Georgette Douwma / Stoplight parrotfish (Sparisoma viride) initial phase. Bonaire, Netherlands Antilles, Caribbean, Atlantic Ocean.
Stoplight parrotfish (Sparisoma viride) terminal phase.  Bonaire, Netherlands Antilles, Caribbean, Atlantic Ocean.
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Georgette Douwma / Stoplight parrotfish (Sparisoma viride) terminal phase. Bonaire, Netherlands Antilles, Caribbean, Atlantic Ocean.
Bumphead parrotfish (Bolbometopon muricatum) shoal. Sipadan Island, Sabah, Borneo.
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Juergen Freund / Bumphead parrotfish (Bolbometopon muricatum) shoal. Sipadan Island, Sabah, Borneo.
Parrotfish (Scaridae) for sale, Suva Seafood Market, Viti Levu, Fiji, South Pacific, April 2014.
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Pete Oxford / Parrotfish (Scaridae) for sale, Suva Seafood Market, Viti Levu, Fiji, South Pacific, April 2014.
Group of Humphead parrotfish (Bolbometopon muricatum) swimming above coral reef,  Sudan. Red Sea.
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Pascal Kobeh / Group of Humphead parrotfish (Bolbometopon muricatum) swimming above coral reef, Sudan. Red Sea.
Two Humphead parrotfish (Bolbometopon muricatum) swimming on above coral reef,  Sudan. Red Sea.
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Pascal Kobeh / Two Humphead parrotfish (Bolbometopon muricatum) swimming on above coral reef, Sudan. Red Sea.
Humphead parrotfish (Bolbometopon muricatum) above coral reef, cleaned by two Blue streaked cleaner wrasses (Labroides dimidiatus) among Jewel fairy basslets / anthias (Pseudanthias squamipinnis) Suda...
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Pascal Kobeh / Humphead parrotfish (Bolbometopon muricatum) above coral reef, cleaned by two Blue streaked cleaner wrasses (Labroides dimidiatus) among Jewel fairy basslets / anthias (Pseudanthias squamipinnis) Suda...
Humphead parrotfish (Bolbometopon muricatum) swimming in open water, Sudan. Red Sea.
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Pascal Kobeh / Humphead parrotfish (Bolbometopon muricatum) swimming in open water, Sudan. Red Sea.
Group of Humphead parrotfish (Bolbometopon muricatum) swimming by coral drop off with divers in the background,  Sudan. Red Sea. June 2013.
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Pascal Kobeh / Group of Humphead parrotfish (Bolbometopon muricatum) swimming by coral drop off with divers in the background, Sudan. Red Sea. June 2013.
Rusty parrotfish (Scarus ferrugineus) close up of tail, coast of Dhofar and Hallaniyat islands, Oman. Arabian Sea.
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Pascal Kobeh / Rusty parrotfish (Scarus ferrugineus) close up of tail, coast of Dhofar and Hallaniyat islands, Oman. Arabian Sea.
Close-up of the head of Bullethead parrotfish (Chlorurus sordidus) sleeping at night in its mucus cocoon, Great Barrier Reef, Australia. Coral sea.
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Pascal Kobeh / Close-up of the head of Bullethead parrotfish (Chlorurus sordidus) sleeping at night in its mucus cocoon, Great Barrier Reef, Australia. Coral sea.
Close-up of the head of a Bullethead parrotfish (Chlorurus sordidus) sleeping at night, Great Barrier Reef, Australia. Coral sea.
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Pascal Kobeh / Close-up of the head of a Bullethead parrotfish (Chlorurus sordidus) sleeping at night, Great Barrier Reef, Australia. Coral sea.
The detail of an eye of Bullethead parotfish (Chlorurus sordidus) taken at night. Gubal Barge Wreck, Gubal Island, Egypt. Strait Of Gubal, Egypt.
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Alex Mustard / The detail of an eye of Bullethead parotfish (Chlorurus sordidus) taken at night. Gubal Barge Wreck, Gubal Island, Egypt. Strait Of Gubal, Egypt.
Parrotfish (Scarus sp) fin detail, Red Sea, Egypt
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Chris Newbert / Parrotfish (Scarus sp) fin detail, Red Sea, Egypt
Parrotfish (Scarus sp) close-up of eye, Red Sea, Egypt
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Chris Newbert / Parrotfish (Scarus sp) close-up of eye, Red Sea, Egypt
Dark-capped Parrotfish (Scarus oviceps) pectoral fin detail, Solomon Islands
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Chris Newbert / Dark-capped Parrotfish (Scarus oviceps) pectoral fin detail, Solomon Islands
Quoy's parrotfish (Scarus quoyi) head portrait, Omaha's Henry Doorly Zoo & Aquarium. Captive, occurs in western Pacific Ocean.
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Joel Sartore / Photo Ark / Quoy's parrotfish (Scarus quoyi) head portrait, Omaha's Henry Doorly Zoo & Aquarium. Captive, occurs in western Pacific Ocean.
Quoy's parrotfish (Scarus quoyi) portrait, Omaha's Henry Doorly Zoo & Aquarium. Captive, occurs in western Pacific Ocean.
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Joel Sartore / Photo Ark / Quoy's parrotfish (Scarus quoyi) portrait, Omaha's Henry Doorly Zoo & Aquarium. Captive, occurs in western Pacific Ocean.
Quoy's parrotfish (Scarus quoyi) portrait, Omaha's Henry Doorly Zoo & Aquarium. Captive, occurs in western Pacific Ocean.
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Joel Sartore / Photo Ark / Quoy's parrotfish (Scarus quoyi) portrait, Omaha's Henry Doorly Zoo & Aquarium. Captive, occurs in western Pacific Ocean.
Quoy's parrotfish (Scarus quoyi) dorsal fin detail, Omaha's Henry Doorly Zoo & Aquarium. Captive, occurs in western Pacific Ocean.
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Joel Sartore / Photo Ark / Quoy's parrotfish (Scarus quoyi) dorsal fin detail, Omaha's Henry Doorly Zoo & Aquarium. Captive, occurs in western Pacific Ocean.
Gulf parrotfish (Scarus persicus) portrait, Sharjah Aquarium UAE. Captive.
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Joel Sartore / Photo Ark / Gulf parrotfish (Scarus persicus) portrait, Sharjah Aquarium UAE. Captive.
Gulf parrotfish (Scarus persicus) portrait, Sharjah Aquarium UAE. Captive.
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Joel Sartore / Photo Ark / Gulf parrotfish (Scarus persicus) portrait, Sharjah Aquarium UAE. Captive.
Green-face parrotfish (Scarus prasiognathos) shoal working together to invade the feeding ground of Lined surgeonfish (Acanthurus lineatus) on a coral reef, Raja Ampat, West Papua, Ceram Sea, Pacific...
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Alex Mustard / Green-face parrotfish (Scarus prasiognathos) shoal working together to invade the feeding ground of Lined surgeonfish (Acanthurus lineatus) on a coral reef, Raja Ampat, West Papua, Ceram Sea, Pacific...
Stareye parrotfish (Calotomus carolinus) terminal phase male, swimming over reef, Hawaii, Pacific Ocean.
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David Fleetham / Stareye parrotfish (Calotomus carolinus) terminal phase male, swimming over reef, Hawaii, Pacific Ocean.
Green sea turtle (Chelonia mydas) being cleaned of algae by Yellow tangs (Zebrasoma flavescens), Chevron tangs (Ctenochaetus hawaiiensis), Black durgon triggerfish (Melichthys niger) and Parrotfish (S...
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Doug Perrine / Green sea turtle (Chelonia mydas) being cleaned of algae by Yellow tangs (Zebrasoma flavescens), Chevron tangs (Ctenochaetus hawaiiensis), Black durgon triggerfish (Melichthys niger) and Parrotfish (S...
Reef manta (Mobula alfredi) getting cleaned of parasites, dead skin, bacteria, and mucus by Blacklip butterflyfish (Chaetodon kleinii) and Moon wrasse (Thalassoma lunare) at a cleaning station in Raja...
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Russell Laman / Reef manta (Mobula alfredi) getting cleaned of parasites, dead skin, bacteria, and mucus by Blacklip butterflyfish (Chaetodon kleinii) and Moon wrasse (Thalassoma lunare) at a cleaning station in Raja...
Bumphead parrotfish (Bolbometopon muricatum) and King angelfish (Holacanthus passer) swimming over reef, Gordon Rocks, Galapagos National Park, Pacific Ocean.
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Lucas Bustamante / Bumphead parrotfish (Bolbometopon muricatum) and King angelfish (Holacanthus passer) swimming over reef, Gordon Rocks, Galapagos National Park, Pacific Ocean.
Roundhead parrotfish (Chlorurus strongycephalus) portrait, South Ari Atoll, Maldives, Indian Ocean.
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Alex Mustard / Roundhead parrotfish (Chlorurus strongycephalus) portrait, South Ari Atoll, Maldives, Indian Ocean.
Red Sea steephead parrotfish (Chlorurus gibbus) male, portrait, Ras Mohammed National Park, Sinai, Egypt, Red Sea.
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Alex Mustard / Red Sea steephead parrotfish (Chlorurus gibbus) male, portrait, Ras Mohammed National Park, Sinai, Egypt, Red Sea.
Bumphead parrotfish (Bolbometopon muricatum) grinding coral / feeding off  Sipadan Island, Malaysia, Celebes Sea, Pacific Ocean.
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David Fleetham / Bumphead parrotfish (Bolbometopon muricatum) grinding coral / feeding off Sipadan Island, Malaysia, Celebes Sea, Pacific Ocean.
Male Bullethead parrotfish (Chlorurus sordidus) fin detail, Sharm El Sheikh, Egypt, Red Sea.
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Alex Mustard / Male Bullethead parrotfish (Chlorurus sordidus) fin detail, Sharm El Sheikh, Egypt, Red Sea.
Male Bullethead parrotfish (Chlorurus sordidus) eye detail, Sharm El Sheikh, Egypt, Red Sea.
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Alex Mustard / Male Bullethead parrotfish (Chlorurus sordidus) eye detail, Sharm El Sheikh, Egypt, Red Sea.
Two male Bullethead parrotfish (Chlorurus sordidus), one with a Cave cleaner shrimp (Urocaridella sp) on its nose, sleeping in crevices in a coral reef at night.  Sharm El Sheikh, Egypt, Red Sea.
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Alex Mustard / Two male Bullethead parrotfish (Chlorurus sordidus), one with a Cave cleaner shrimp (Urocaridella sp) on its nose, sleeping in crevices in a coral reef at night. Sharm El Sheikh, Egypt, Red Sea.
Male Bullethead parrotfish (Chlorurus sordidus) with a Cave cleaner shrimp (Urocaridella sp.) on its head, sleeping in a crevice in a coral reef, at night,  Sharm El Sheikh, Egypt, Red Sea.
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Alex Mustard / Male Bullethead parrotfish (Chlorurus sordidus) with a Cave cleaner shrimp (Urocaridella sp.) on its head, sleeping in a crevice in a coral reef, at night, Sharm El Sheikh, Egypt, Red Sea.
RF - Composite image of blue tropical reef fish on a black background, Blue triggerfish (Pseudobalistes fuscus), Elonggate surgeonfish (Acanthurus mata), Yellowtail tang (Zebrasoma xanthurum), Neon da...
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Georgette Douwma / RF - Composite image of blue tropical reef fish on a black background, Blue triggerfish (Pseudobalistes fuscus), Elonggate surgeonfish (Acanthurus mata), Yellowtail tang (Zebrasoma xanthurum), Neon da...
RF - Composite image of tropical reef fish on a black background, Rusty parrotfish (Scarus ferrugineus), Striped surgeonfish (Acanthurus lineatus), Saddleback butterflyfish (Chaetodon falcula), Seahor...
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Georgette Douwma / RF - Composite image of tropical reef fish on a black background, Rusty parrotfish (Scarus ferrugineus), Striped surgeonfish (Acanthurus lineatus), Saddleback butterflyfish (Chaetodon falcula), Seahor...
Tropical reef fish composite image on black background, Kasmira snapper (Lutjanus kasmira), Greenthroat parrotfish (Scarus prasiognathos), Two spot wrasse (Oxycheilinus bimaculatus), Rusty parrotfish...
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Georgette Douwma / Tropical reef fish composite image on black background, Kasmira snapper (Lutjanus kasmira), Greenthroat parrotfish (Scarus prasiognathos), Two spot wrasse (Oxycheilinus bimaculatus), Rusty parrotfish...
Blue tropical reef fish composite image on black background, Blue triggerfish (Pseudobalistes fuscus), Bicolour angelfish (Centropyge bicolor), Elongate surgeonfish (Acanthurus mata), Yellowtail tang...
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Georgette Douwma / Blue tropical reef fish composite image on black background, Blue triggerfish (Pseudobalistes fuscus), Bicolour angelfish (Centropyge bicolor), Elongate surgeonfish (Acanthurus mata), Yellowtail tang...
Two Stoplight parrotfish (Sparisoma viride), showing terminal phase and initial phase variations, composite image on black background, Bonaire, Netherlands Antilles, Caribbean.
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Georgette Douwma / Two Stoplight parrotfish (Sparisoma viride), showing terminal phase and initial phase variations, composite image on black background, Bonaire, Netherlands Antilles, Caribbean.
Greenthroat / Singapore parrotfish (Scarus prasiognathos), male and two females, composite image on black background, Andaman Sea, Thailand,
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Georgette Douwma / Greenthroat / Singapore parrotfish (Scarus prasiognathos), male and two females, composite image on black background, Andaman Sea, Thailand,
Humphead parrotfish (Bolbometopon muricatum) showing breeding colouration, spawning in the early morning, Palau, Micronesia, Pacific Ocean.
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Henley Spiers / Humphead parrotfish (Bolbometopon muricatum) showing breeding colouration, spawning in the early morning, Palau, Micronesia, Pacific Ocean.
Bullethead parrotfish (Chlorurus sordidus) sleeping at night in its mucus cocoon, Red Sea, Egypt.
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Pascal Kobeh / Bullethead parrotfish (Chlorurus sordidus) sleeping at night in its mucus cocoon, Red Sea, Egypt.
Stoplight parrotfish (Sparisoma viride) sleeping at night, The Gardens of the Queen, Cuba, Caribbean Sea.
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Pascal Kobeh / Stoplight parrotfish (Sparisoma viride) sleeping at night, The Gardens of the Queen, Cuba, Caribbean Sea.
Stoplight parrotfish (Sparisoma viride) swimming towards reef and feeding on coral, Bonaire, Leeward Antilles, Caribbean.
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Solvin Zankl / Stoplight parrotfish (Sparisoma viride) swimming towards reef and feeding on coral, Bonaire, Leeward Antilles, Caribbean.
Colourful coral reef wall, with orange scalefin anthias (Pseudanthias squamipinnis) swarming over red, orange and pink soft corals (Dendronephthya sp.) in a current, with a lone bullethead parrotfish...
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Alex Mustard / Colourful coral reef wall, with orange scalefin anthias (Pseudanthias squamipinnis) swarming over red, orange and pink soft corals (Dendronephthya sp.) in a current, with a lone bullethead parrotfish...