Anup Shah / Eastern Chimpanzee (Pan troglodytes schweinfurthii) forty-three year old female, named Gremlin, scratching face, Gombe National Park, Tanzania

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 41 years grooms her daughter 'Golden' aged 14 years while their infants 'Gizmo' aged 3 years and 'Glamma' aged 9 months sit b...

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 40 years holding her daughter 'Glitter's' baby daught aged 2 months and her own infant son aged 2 years. With 'Glitter' aged...

Fiona Rogers / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 40 years holding female baby aged 2, her granddaughter. Gombe National Park, Tanzania. Gremlin took the new born baby from he...

Fiona Rogers / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 40 years calling while holding her baby granddaughter aged 2 months and her own infant son aged 2 years. Gombe National Park,...

Anup Shah / Young Chimpanzees + mother fishing for termites, Gombe NP, Tanzania 2003 'Gremlin'

Anup Shah / Chimpanzee + young in tree 'Gremlin' + 'G', Gombe NP, Tanzania 2003

Anup Shah / Male Chimpanzee + young fishing for termites, Gombe NP, Tanzania 2003 'Gremlin'

Anup Shah / Female Chimpanzee 'Gremlin' 31-years-old, carrying twin daughters Golden and Glitter, 3-years-3-monhts-old, jockey riding {Pan troglodytes schweinfurtheii} Gombe NP, Tanzania. 2002

Anup Shah / Young male Chimpanzee 'Gold' 3-years-3-months-old, watches mother 'Gremlin' use stick as tool to catch termites {Pan troglodytes schweinfurtheii} Gombe NP, Tanzania. 2002

Anup Shah / Female Chimpanzee 'Gremlin' demonstrates to offspring how to use a stem as a tool to remove termites from a termite mound {Pan troglodytes schweinfurtheii} Kasekela community, Gombe NP, Tanzania. 2002

Anup Shah / Chimpanzee mother 'Gremlin' carries female twins 'Golden' (in front) and 'Glitta' (behind) {Pan troglodytes schweinfurtheii} Kasekela community, Gombe NP, Tanzania. 2002

Anup Shah / Female Chimpanzee 'Gremlin' carries female twins 'Golden' and 'Glitta' {Pan troglodytes schweinfurtheii} Kasekela community, Gombe NP, Tanzania. 2002

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 43 years and her sons 'Gimli' aged 10 years and 'Gizmo' aged 4 years . Gombe National Park, Tanzania....

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 43 years walking . Gombe National Park, Tanzania. September 2014.

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 43 years feeding on charcoal . Gombe National Park, Tanzania. September 2014.

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 42 years carrying her infant son 'Gizmo' aged 4 years on her back.Gombe National Park, Tanzania.

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 42 years playing with her grandaughter 'Glamour' aged 2 years.Gombe National Park, Tanzania. September 2013.

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 42 years playing with her grandaughter 'Glamour' aged 2 years.Gombe National Park, Tanzania. September 2013.

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 42 years being groomed by her daughter 'Golden' aged 15 years.Gombe National Park, Tanzania. September 2013.

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 42 years walking along a trail followed by her family.Gombe National Park, Tanzania. September 2013.

Anup Shah / Eastern Chimpanzee (Pan troglodytes schweinfurthii) forty-two year old female, named Gremlin, playing with her two year old granddaughter, named Glamour, Gombe National Park, Tanzania

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 42 years sharing figs with her juvenile son 'Gizmo' aged 3 years and 9 months . Gombe National Park, Tanzania.

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 42 years and her juvenile son 'Gizmo' aged 3 years and 9 months feeding on figs . Gombe National Park, Tanzania.

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 42 years carrying her son 'Gizmo' aged 3 years and 9 months on her back . Gombe National Park, Tanzania.

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 42 years feeding on figs . Gombe National Park, Tanzania.

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'gremlin' aged 41 years carrying her infant son 'Gizmo' aged 3 years walking along a forest trail. Gombe National Park, Tanzania. October 20...

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 41 years carrying her infant son 'Gizmo' age 3 years sitting in bushes. Gombe National Park, Tanzania. May 2012.

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 41 years and her daughter 'Gaia' aged 19 years termite fishing while their sons play. Gombe National Park, Tanzania. May 2012...

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 41 years head and shoulders portrait. Gombe National Park, Tanzania. May 2012.

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) 'Gimli' aged 8 years grooming his mother 'Gremlin' aged 41 years. Gombe National Park, Tanzania. May 2012.

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 41 years and infant son 'Gizmo' aged 3 years climbing down a tree. Gombe National Park, Tanzania. May 2012.

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 41 years carrying her infant son 'Gizmo' aged 3 years on her back. Gombe National Park, Tanzania. May 2012.

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 41 years feeding on tabogoro fruit with her daughter 'Golden' aged 14 years and her son 'Gimli' aged 8 years. Gombe National...

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 40 years callng, while holding her daughter 'Glitter's' baby aged 3 months and her own infant son 'Gizmo' aged 2 years. Gombe...

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 40 years holding her daughter 'Glitter's' baby daughter aged 2 months. Gombe National Park, Tanzania. Gremlin took the new bo...

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) male infant 'Gizmo' aged 2 years suckling from his mother 'Gremlin' next to the female baby of 'Glitter' aged 2 months. Gombe National Park, Tanzan...

Fiona Rogers / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female baby aged 2 months, the baby of adolescent female 'Glitter' held by her grandmother 'Gremlin' aged 40 years. Gombe National Park, Tanzania....

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 40 years climbing a branch with her daughter 'Glitter's' baby age 2 months and her own infant son 'Gizmo' aged 2 years. Gombe...

Fiona Rogers / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 40 years holding her daughter 'Glitter's' first born daughter aged 2 months. Gombe National Park, Tanzania. Gremlin took the...

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 40 years holding her daughter 'Glitter's' baby aged 2 months and her infant son aged 2 years. Gombe National Park, Tanzania....

Anup Shah / Olive baboon female (Papio cynocephalus anubis) presenting to Eastern chimpanzee (Pan troglodytes schweinfurtheii) juvenile male 'Gimli' aged 7 years watched by his mother 'Gremlin'. Gombe National Pa...

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 40 years holding her daughter 'Glitter's' baby aged 2 months. Gombe National Park, Tanzania. Gremlin took the new born baby f...

Fiona Rogers / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 40 years holding her daughter 'Glitter's' baby daughter aged 2 months. With 'Glitter' aged 13 years watching something that h...

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 40 years holding her daughter 'Glitter's' baby daughter aged 2 months. Gombe National Park, Tanzania. Gremlin took the new bo...

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 40 years holding her daughter 'Glitter's' baby daughter aged 2 months and her own infant son aged 2 years. With 'Glitter' age...

Fiona Rogers / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female baby aged 2 months, the baby of adolescent female 'Glitter' held by her grandmother 'Gremlin' aged 40 years. Gombe National Park, Tanzania....

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 40 years head and shoulders portrait. Gombe National Park, Tanzania.

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 40 years head and shoulders portrait. Gombe National Park, Tanzania.

Fiona Rogers / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 40 years, holding her granddaughter, age 2 months. Gombe National Park, Tanzania. Gremlin took the new born baby from her dau...

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 40 years drinking from a stream while carrying her daughter 'Glitter's' first born daughter aged 2 months and her own infant...

Fiona Rogers / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 40 years holding her granddaughter aged 2 months. Gombe National Park, Tanzania. Gremlin took the new born baby from her daug...

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 40 years holding her daughter 'Glitter's' baby aged 3 months. With her daughter 'Gaia' aged 18 years and infant son 'Gizmo' a...

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 40 years walking while carrying her daughter 'Glitter's' baby aged 2 months and her own infant son 'Gizmo' aged 2 years. Gomb...

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 40 years holding her daughter 'Glitter's' baby aged 3 months while her infant son 'Gizmo' aged 2 years sits alongside. Gombe...

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 40 years with male infant 'Gizmo' aged 2 years and the female baby of 'Glitter' aged 2 months suckling. Gombe National Park,...

Fiona Rogers / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 40 years being groomed by her daughter 'Gaia' aged 18 years. Gombe National Park, Tanzania.

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 40 years with her infant son 'Gizmo' aged 1-2 years sitting between two trees. Gombe National Park, Tanzania.

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 40 years with her infant son 'Gizmo' aged 1-2 years sitting between two trees. Gombe National Park, Tanzania.

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 40 years picking her nose and eating it. Gombe National Park, Tanzania.

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 40 years feeding on Mbula fruit. Gombe National Park, Tanzania.

Anup Shah / Eastern Chimpanzee (Pan troglodytes schweinfurthii) forty year old female, named Gremlin, carrying young and scanning canopy for fruit, with her seven year old juvenile son, named Gimli, in the foregr...

Anup Shah / Chimpanzee eating termites off twig, Gombe NP, Tanzania 2003 'Gremlin'

Anup Shah / Young Chimpanzee jockey riding on mother, Gombe NP, Tanzania 2003 'Gremlin'

Anup Shah / Chimpanzee + young in tree, 'Gremlin' + 'G', Gombe NP, Tanzania 2003

Anup Shah / Male Chimpanzee eating termites off twig tool, Gombe NP, Tanzania 2003 'Gremlin'

Anup Shah / Male Chimpanzee tool making at termite mound, 'Gremlin' Gombe NP, Tanzania 2003

Anup Shah / Male Chimpanzee tool making at termite mound, 'Gremlin' Gombe NP, Tanzania 2003

Anup Shah / Chimpanzee grooming young 'Gremlin' + 'G', Gombe NP, Tanzania 2002

Anup Shah / Chimpanzee suckling young, Gombe NP, Tanzania, 'Gremlin' + 'G' 2002

Anup Shah / Chimpanzee carrying twins, Gombe NP, Tanzania. Gremlin with twin daughters Gold and Glitter

Anup Shah / Chimpanzee grasping vegetation with foot, 'Gremlin' female 31-years-old, {Pan troglodytes schweinfurtheii} Gombe NP, Tanzania. 2002

Anup Shah / Female Chimpanzee 'Gremlin' 31-years-old, {Pan troglodytes schweinfurtheii} Gombe NP, Tanzania. 2002

Anup Shah / Female Chimpanzee 'Gremlin' demonstrates to offspring how to use a stem as a tool to remove termites from a termite mound {Pan troglodytes schweinfurtheii} Kasekela community, Gombe NP, Tanzania. 2002

Anup Shah / Eastern Chimpanzee (Pan troglodytes schweinfurthii) forty-two year old female, named Gremlin, carrying her four year old son, named Gizmo, Gombe National Park, Tanzania

Anup Shah / Eastern Chimpanzee (Pan troglodytes schweinfurthii) forty-two year old female, named Gremlin, carrying her four year old son, named Gizmo, Gombe National Park, Tanzania

Anup Shah / Eastern Chimpanzee (Pan troglodytes schweinfurthii) forty year old female, named Gremlin, scanning canopy for fruit, Gombe National Park, Tanzania

Anup Shah / Eastern Chimpanzee (Pan troglodytes schweinfurthii) forty year old female, named Gremlin, carrying two year old infant, named Gizmo, Gombe National Park, Tanzania