Florian Möllers / Golden snub-nosed monkeys (Rhinopithecus roxellana qinlingensis) family group / harem resting / sleeping / cuddling, Zhouzhi Nature Reserve, Qinling mountains, Shaanxi, China, April 2006

Yva Momatiuk & John Eastcott / Mustang (Equus caballus) harem stallions posturing as prelude to fight, Montana

Miles Barton / Gelada baboons, harem male with female + young {Theropithecus gelada}, Simien Mtns, Ethiopia,

XI ZHINONG / Tibetan antelope (Pantholops hodgsoni) male running after his harem of females, Kekexili, Qinghai, December

Christophe Courteau / Herd of Impalas (Aepyceros melampris) during the rutting season. A harem with one male and several females, Botswana. Buffalo in background

Christophe Courteau / Male impala running to gather the females in his harem {Aepyceros melampus}, Botswana.

Anup Shah / Mature male gelada (Theropithecus gelada) looks up from amongst his harem of females. Simien Mountains National Park, Ethiopia. Nov 2008.

George Sanker / Elk / Wapiti (Cervus canadensis) bull with his harem of females. Yellowstone National Park, Wyoming, USA, October.

George Sanker / Elk / Wapiti (Cervus canadensis) bull with harem by the Gardiner River. Yellowstone National Park, Wyoming, USA, September.

Charlie Summers / Steller Sea Lion (Eumetopias jubatus) 'beachmaster' or dominant male with his harem gathered round on the rocks of West Brothers Island. Frederick Sound, Alaska, July.

Hiroya Minakuchi / Steller's Sea Lion (Eumetopias jubatus) bull and harem, Prince William Sound, Alaska

Wild Wonders of Europe / Lesniewski / Red Deer (Cervus elaphus) females, a harem belonging to an alpha male. Bieszczady, Carpathian Mountains, Poland, September.

Brent Stephenson / Large adult male Steller's Sea Lion (Eumetopias jubatus) resting on rocks surrounded by females of his harem. Inian Islands, Alaska, United States, July.

Florian Möllers / Golden snub-nosed monkey (Rhinopithecus roxellana qinlingensis) family group / harem resting / grooming in a tree, Zhouzi Nature Reserve, Qinling mountains, Shaanxi, China. April 2006

Florian Möllers / Golden snub-nosed monkeys (Rhinopithecus roxellana qinlingensis) family group / harem resting in tree, Zhouzhi Nature Reserve, Qinling mountains, Shaanxi, China, April 2006

Florian Möllers / Golden snub-nosed monkey (Rhinopithecus roxellana qinlingensis) family group / harem moving through trees, Zhouzhi Nature Reserve, Qinling mountains, Shaanxi, China, April 2006

Florian Möllers / Golden snub-nosed monkey (Rhinopithecus roxellana qinlingensis) family group / harem sleeping in a tree, Zhouzi Nature Reserve, Qinling mountains, Shaanxi, China. April 2006

Florian Möllers / Golden snub-nosed monkey (Rhinopithecus roxellana qinlingensis) family group / harem resting / grooming in a tree, Zhouzhi, Shaanxi, China.

Florian Möllers / Golden snub-nosed monkeys (Rhinopithecus roxellana qinlingensis) family group / harem resting / sleeping / cuddling, Zhouzhi Nature Reserve, Qinling mountains, Shaanxi, China, April 2006

Florian Möllers / Golden snub-nosed monkey (Rhinopithecus roxellana qinlingensis) family group / harem resting / grooming in a tree, Zhouzi Nature Reserve, Qinling mountains, Shaanxi, China. April 2006

Florian Möllers / Golden snub-nosed monkeys (Rhinopithecus roxellana qinlingensis) family group /harem resting in tree, Zhouzhi Nature Reserve, Qinling mountains, Shaanxi, China, April 2006

Frederique Olivier / Aerial view of Southern elephant seal (Mirounga leonina) harem on beach, Macquarie Island, Southern Atlantic, Australian Antarctica, November 2007

Pete Oxford / Bull Southern Elephant Seal (Mirounga leonina) after fighting for take over of the harem. Even a short fight usually results in both bulls getting badly cut on their necks and faces. The Tussock Bird...

Pete Oxford / Southern Elephant Seal (Mirounga leonina) Bull vocalising after fighting for control of the harem. Sea Lion Island. South of mainland east Falkland Island. Falkland Islands

Pete Oxford / Southern Elephant Seal (Mirounga leonina) Bull after fighting for control of the harem. Sea Lion Island. South of mainland east Falkland Island. Falkland Islands

Pete Oxford / Colony / Harem of Southern Elephant Seals (Mirounga leonina) lying on beach, Sea Lion Island. South of mainland east Falkland Island. Falkland Islands

Suzi Eszterhas / Steller's Sea Lion (Eumetopias jubatus) male with small harem calling, Katmai National Park, Alaska

LYNN M. STONE / Wild horse / Mustang (Equus caballus) grey stallion 'Cloud' interacting with two mares from his harem, Pryor Mountains, Wyoming / Montana border, USA

GEORGETTE DOUWMA / Dotted wrasse (Cirrhilabrus punctatus) territorial male with harem females, Papua New Guinea.

GEORGETTE DOUWMA / Dotted wrasse (Cirrhilabrus punctatus) male with his harem of females, Papua New Guinea

Kristel Richard / Wild Przewalski (or Takhi) horse {Equus ferus przewalski} endangered species, breeding stallion herding his harem of females, Hustai National Park, Tuv Province, Mongolia, July 2007

Anup Shah / Harem of female Gelada baboons (Theropithecus gelada) confronting a rival male, aggression shown with lip flips, Simien Mountains National Park, Ethiopia, November

Matthias Breiter / Steller's Sea Lion (Eumetopias jubatus) male and harem of females on rock, Shelikof Strait, Alaska

Jim Brandenburg / Steller's Sea Lion (Eumetopias jubatus) male and females, Prince William Sound, Alaska

Donald M. Jones / Mule Deer (Odocoileus hemionus) does and fawns with a buck, western Montana

Donald M. Jones / Mule Deer (Odocoileus hemionus) does and fawns with a buck, western Montana

Donald M. Jones / Mule Deer (Odocoileus hemionus) buck with does during rut, western Montana

Donald M. Jones / American Elk (Cervus elaphus nelsoni) bull and cows in winter, western Montana

Donald M. Jones / American Elk (Cervus elaphus nelsoni) bull bugling with harem, western Alberta, Canada

Donald M. Jones / American Elk (Cervus elaphus nelsoni) bull and cows, western Alberta, Canada

GEORGETTE DOUWMA / Lyretail anthias / Goldie (Pseudanthias squamipinnis) male with 'harem' of females. Red Sea, Egypt.

Matthias Breiter / Northern Fur Seal (Callorhinus ursinus) bull and harem, Pribilof Islands, Alaska

Roy Mangersnes / Southern elephant seal (Mirounga leonina) male with his harem of females on the beach, Moltke Harbour, South Georgia, Antarctica. November

Roy Mangersnes / Southern elephant seal (Mirounga leonina) male protecting his harem of females on the beach, Moltke Harbour, South Georgia, Antarctica. Cub is being flattened. November, sequence 2/2

Roy Mangersnes / Southern elephant seal (Mirounga leonina) harem with females and cubs, Fortuna Bay, South Georgia, Antarctica. November

Roy Mangersnes / Southern elephant seal (Mirounga leonina) male protecting his harem of females on the beach, Moltke Harbour, South Georgia, Antarctica. Cub is about to be flattened. November, sequence 1/2

Roy Mangersnes / Southern elephant seal (Mirounga leonina) pair mating. Satellite male mounting a harem female before the dominant male beach master realizes. Fortuna Bay, South Georgia, Antarctica. November

Yva Momatiuk & John Eastcott / Southern Elephant Seal (Mirounga leonina) bull waiting in surf near harem to intercept females during breeding season, St. Andrews Bay, South Georgia Island

Tim Fitzharris / Impala (Aepyceros melampus), buck and harem on savanna, Kenya

Michio Hoshino / Dall's Sheep (Ovis dalli) male with harem, Alaska

Yva Momatiuk & John Eastcott / Southern Elephant Seal (Mirounga leonina) harem during breeding season in spring, with snowy Allardyce Range in background, St. Andrews Bay, South Georgia Island

Yva Momatiuk & John Eastcott / King Penguin (Aptenodytes patagonicus) group walking among harems of Southern Elephant Seals (Mirounga leonina) on beach, St. Andrews Bay, South Georgia Island

Yva Momatiuk & John Eastcott / Southern Elephant Seal (Mirounga leonina) bull mating with female on beach outside harem, St. Andrews Bay, South Georgia Island

Yva Momatiuk & John Eastcott / Southern Elephant Seal (Mirounga leonina) female and pup resting in harem on beach during breeding season, St. Andrews Bay, South Georgia Island

Yva Momatiuk & John Eastcott / Southern Elephant Seal (Mirounga leonina) harem on beach during breeding season with Graae Glacier and snowy mountains in background, Trollhul, South Georgia Island

Yva Momatiuk & John Eastcott / Southern Elephant Seal (Mirounga leonina) female and pup in harem, Trollhul, South Georgia Island

Yva Momatiuk & John Eastcott / Southern Elephant Seal (Mirounga leonina) harem on beach during pupping and breeding season in spring, mountains of Allardyce Range behind, South Georgia Island

Yva Momatiuk & John Eastcott / Southern Elephant Seal (Mirounga leonina) bull and his harem of females and young black pups resting on beach during breeding season, South Georgia Island

George Chan / A male Tibetan antelope / Chiru (Pantholops hodgsonii) guarding his female harem in the Chang Tang nature reserve of central Tibet during the rutting season. December 2006

George Chan / Sub-adult Tibetan antelope / Chiru (Pantholops hodgsonii) in the Chang Tang Nature Reserve of central Tibet. During the rutting season adult males gather females in a harem while juveniles wander the...

George Chan / Male Tibetan antelope / Chiru (Pantholops hodgsonii) chasing his female harem across the Chang Tang Nature Reserve of central Tibet, during the rutting season. December 2006

George Chan / Sub-adult Tibetan antelope / Chiru (Pantholops hodgsonii) in the Chang Tang Nature Reserve of central Tibet. During the rutting season adult males gather females in a harem while juveniles wander the...

George Chan / Male Tibetan antelope / Chiru (Pantholops hodgsonii) guarding his female harem in the Chang Tang Nature Reserve of central Tibet, bellowing load calls to warn off other males. December 2006,

George Chan / A male Tibetan antelope / Chiru (Pantholops hodgsonii) guarding his female harem in the Chang Tang nature reserve of central Tibet during the rutting season. December 2006

George Chan / Sub-adult Tibetan antelope / Chiru (Pantholops hodgsonii) in the Chang Tang Nature Reserve of central Tibet. During the rutting season adult males gather females in a harem while juveniles wander the...

George Chan / Sub-adult Tibetan antelope / Chiru (Pantholops hodgsonii) in the Chang Tang Nature Reserve of central Tibet. During the rutting season adult males gather females in a harem while juveniles wander the...

George Chan / A male Tibetan antelope / Chiru (Pantholops hodgsonii) guarding his female harem in the Chang Tang nature reserve of central Tibet during the rutting season. December 2006

Yva Momatiuk & John Eastcott / Pronghorn Antelope (Antilocapra americana) buck and harem on grassland, fall, Wyoming

Suzi Eszterhas / Antarctic Fur Seal (Arctocephalus gazella) colony, Hercules Bay, South Georgia

Elio Della Ferrera / Male Hamadryas Baboon (Papio hamadryas) with his harem of females and their young, Ethiopia 1992

Elio Della Ferrera / Male Hamadryas Baboon (Papio hamadryas) with his harem of females and young, Ethiopia 1992

Flip Nicklin / Steller's Sea Lion (Eumetopias jubatus) male with harem of females and pups, Alaska

JIM CLARE / Greater spear nosed bat harem {Phyllostomus hastatus} limestone cave, Trinidad

Jabruson / Uganda kob pair mating {Kobus kob thomasi} Virunga NP, Democratic Republic of Congo, formerly Zaire

Martin Dohrn / Impala (Aepyceros melampus) rear view of a male following female herd, Masai Mara GR, Kenya

Ian McCarthy / Male Southern elephant seal with harem {Mirounga leonina} Marion Island, sub-antarctica

Shin Yoshino / Elk (Cervus elaphus) buck and two does at sunset, Yellowstone National Park, Wyoming

Mark Raycroft / White-tailed Deer (Odocoileus virginianus) ten point buck in meadow with grazing does

Mark Raycroft / White-tailed Deer (Odocoileus virginianus) buck watching over does in meadow

Mark Raycroft / Elk (Cervus elaphus) large bull and harem of cows crossing railroad tracks in the mountains

Konrad Wothe / Steller's Sea Lion (Eumetopias jubatus) group congregating on rock, West Brothers Island, Inside Passage, Alaska

Konrad Wothe / Steller's Sea Lion (Eumetopias jubatus) group congregating on rock, West Brother Island, Inside Passage, Alaska

Konrad Wothe / Steller's Sea Lion (Eumetopias jubatus) group congregating on rock, West Brothers Island, Inside Passage, Alaska

Yva Momatiuk & John Eastcott / Northern Fur Seal (Callorhinus ursinus) harem group in large rookery, St Paul Island, Pribilof Islands, Alaska

Yva Momatiuk & John Eastcott / Mustang (Equus caballus) harem stallion portrait, autumn, Cayuse Ranch, Wyoming

Michio Hoshino / Alaska Moose (Alces alces gigas) and harem on autumn colored tundra, Alaska

Tui De Roy / Vicuna (Vicugna vicugna) wild Andean camelid, harem holding male on lookout for rivals, Pampa Galeras Nature Reserve, Peru

Mark Moffett / Stalk-eyed Fly (Teleopsis sp) females in a single male's harem gather on rootlets near streams, Malaysia

Michio Hoshino / Alaska Moose (Alces alces gigas) male watches over harem on tundra, Alaska

Michio Hoshino / Alaska Moose (Alces alces gigas) male watches over harem on tundra, Alaska

Michio Hoshino / Alaska Moose (Alces alces gigas) male watches over harem on tundra, Alaska

Pete Oxford / New Zealand / Hooker's sealion male with harem {Phocarctos hookeri} Enderby Is, New Zealand

Pete Oxford / Northern fur seal male and harem {Callorhinus ursinus} Bering Is, Russia Mys

Peter Bassett / Southern elephant seal pair mating {Mirounga leonina} South Georgia.

Pete Oxford / New Zealand sealions mating {Neophoca hookeri} Auckland Is, New Zealand Enderby Island

JIM CLARE / Greater spear nosed bat harem {Phyllostomus hastatus} inside Limestone cave, Trinidad