Tony Heald / Hippopotamus {Hippopotamus amphibius} territorial defense at water edge, Chobe national park, Botswana.

Tony Heald / African Wild Dog (Lycaon pictus) pack passing infront of large African elephant (Loxodonta africana) Mana Pools National Park, Zimbabwe October 2012

Matthew Maran / Wildlife photographer Tony Heald photographing Red deer in Richmond Park, London, UK January 2012. Model released.

Tony Heald / Black faced Impala (Aepyceros melamis petersi) female jumping, Etosha National Park, Namibia

Tony Heald / Black faced Impala (Aepyceros melamis petersi) female jumping, Etosha National Park, Namibia

Tony Heald / African Lion (Panthera leo) young male at sunrise, Etosha National Park, Namibia

Sharon Heald / Common zebra (Equus quagga) foal in dusty sunset, Etosha National Park, Namibia Photo by Sharon Heald

Tony Heald / Black rhinoceros (Diceros bicornis) Etosha National Park, Namibia October

Tony Heald / Black rhinoceros (Diceros bicornis) male urine testing, flehmen reaction, Etosha National Park, Namibia October

Tony Heald / African elephant (Loxodonta africana) at sunset by water, Etosha National Park, Namibia October

Tony Heald / African lion (Panthera leo) young male backlit at sunrise, Etosha National Park, Namibia October

Tony Heald / Eland (Taurotragus oryx) males locking horns and fighting. Devil's Thorn flowers (Tribulus terrestris) cover the ground. Etosha National Park, Namibia, January.

Tony Heald / Black Rhinoceros (Diceros bicornis) running. Etosha National Park, Namibia, January.

Tony Heald / Eland (Taurotragus oryx) male horning the ground which is covered in Devil's Thorn flowers (Tribulus terrestris). Etosha National Park, Namibia, January.

Tony Heald / Spotted Hyaena (Crocuta crocuta) rolling in Devil's Thorn flowers (Tribulus terrestris). Etosha National Park, Namibia, January.

Sharon Heald / Black-faced Impala (Aepyceros melamis petersi) male standing in a field of Devil's Thorn flowers (Tribulus terrestris). Etosha National Park, Namibia, January.

Tony Heald / Black Rhinoceros (Diceros bicornis) charging. Etosha National Park, Namibia, January.

Tony Heald / Red-Billed Teal (Anas erythrorhyncha) foraging in water. Etosha National Park, Namibia, January.

Tony Heald / Zebra (Equus quagga) foal running through Devil's Thorn flowers (Tribulus terrestris). Etosha National Park, Namibia, January.

Tony Heald / Blue Crane (Anthropoides paradiseus) performing its dance display. Etosha National Park, Namibia, January.

Tony Heald / Black-Winged Stilt (Himantopus himantopus) wading. Etosha National Park, Namibia, January.

Tony Heald / African Elephant (Loxodonta africana) male scenting with trunk raised. Etosha National Park, Namibia, January.

Tony Heald / Zebra (Equus quagga) foal peering over its mother's striped back. Monochrome. Etosha National Park, Namibia, January.

Sharon Heald / African lion [Panthera leo] mother leading two very young cubs to water, Etosha National Park, Namibia, August

Tony Heald / White rhinoceros [Ceratotherium simum] at water, Etosha National Park, Namibia, August

Tony Heald / Ostrich [Struthio camelus] courtship display by female, Etosha National Park, Namibia, August

Sharon Heald / African lion [Panthera leo] juvenile males playing, Etosha National Park, Namibia, August

Tony Heald / African elephant [Loxodonta africana] female herd approaching waterhole with trunks raised to smell danger, Etosha National Park, Namibia, August

Tony Heald / African elephant (Loxodonta africana) herd walking to waterhole, Etosha National Park, Namibia, August

Tony Heald / African elephant (Loxodonta africana) at waterhole with Giraffe in the background, Etosha National Park, Namibia, August

Tony Heald / African elephant (Loxodonta africana) at waterhole with Giraffe in the background, Etosha National Park, Namibia, August

Tony Heald / African elephant (Loxodonta africana) threat display by shaking head, at waterhole with Ostrich, Zebra and other wildlife in the background, Etosha National Park, Namibia, August Not available for rin...

Tony Heald / Gemsbok [Oryx gazella] two males fighting, Etosha National Park, Namibia, August

Tony Heald / African lion (Panthera leo) juvenile males play fighting, Etosha National Park, Namibia, August

Tony Heald / African lion (Panthera leo) juvenile males play fighting, Etosha National Park, Namibia, August

Tony Heald / Eland [Taurotragus oryx] male rubbing wet mud onto forehead brush, Etosha National Park, Namibia, August

Tony Heald / Black Rhinoceros [Diceros bicornis] drinking, Etosha National Park, Namibia, August

Tony Heald / Ostrich [Struthio camelus] courtship display by female, Etosha National Park, Namibia, August

Tony Heald / African elephant (Loxodonta africana) at waterhole with trunk in air after spraying itself with water, herd of Zebra drinking in the background, Etosha National Park, Namibia, June

Tony Heald / African elephant (Loxodonta africana) drinking at waterhole with Zebra and Giraffe in background, Etosha National Park, Namibia, June 2009

Tony Heald / African elephant (Loxodonta africana) leaving waterhole with Zebra and Giraffe in the background, Etosha National Park, Namibia, June 2009

Tony Heald / African elephant (Loxodonta africana) in the rain, Etosha National Park, Namibia, January

Tony Heald / Male Eland (Taurotragus oryx) jumping, Etosha National Park, Namibia, January

Tony Heald / Male Eland (Taurotragus oryx) jumping, Etosha National Park, Namibia, January

Tony Heald / Black rhinoceros (Diceros bicornis) charging, Etosha National Park, Namibia, January

Tony Heald / Black rhinoceros (Diceros bicornis) charging, Etosha National Park, Namibia, January

Tony Heald / Black rhinoceros (Diceros bicornis) charging, Etosha National Park, Namibia, January

Tony Heald / Kori bustard (Ardeotis kori) displaying, Etosha National Park, Namibia, January

Tony Heald / Pigtail Macaques (Macaca nemestrina) crossing purpose-built "primate wires" connecting isolated patches of forest, Kinabatangan River, Sukau, Sabah, Borneo, September 2008

Tony Heald / Malayan colugo (Cynocephalus variegatus) hanging upside-down in tree feeding on algae at night, Danum Valley, Sabah, Borneo, September

Tony Heald / Temple / Wagler's Pit Viper [Tropidolaemus wagleri] Bako National Park, Sarawak, Borneo. September

Tony Heald / Long nosed / Oriental Whip Snake [Ahaetulla prasina] Bako National Park, Sarawak, Borneo, September

Tony Heald / Canopy walkway in lowland dipterocarp rainforest, Danum Valley, Sabah, Borneo, September 2008

Tony Heald / Danum River flowing through lowland dipterocarp rainforest with shadow of people on walkway crossing river, Danum Valley, Sabah, Borneo, September 2008

Tony Heald / African Fish Eagle (Haliaeetus vocifer) carrying nesting material, Chobe National Park, Botswana May 2008

Tony Heald / Southern plains grey / Hanuman langur {Semnopithecus dussumieri} baby reaching up to mother's face, Bandhavgarh NP, Madhya Pradesh, India, March

Tony Heald / Bengal tiger [Panthera tigris tigris] crossing road in front of watching tourists, Bandhavgarh NP, Madhya Pradesh, India, March

Sharon Heald / African lion {Panthera leo} male sniffing a scent mark, Etosha, Namibia, January

Tony Heald / African bullfrogs {Pyxicephalus adspersus} group of males in water, Etosha NP, Namibia, January

Tony Heald / Herd of Common zebra {Equus quagga} drinking at waterhole at night, Okaukuejo, Etosha NP, Namibia, January

Tony Heald / African bullfrog {Pyxicephalus adspersus} vanquished male leaps away from fight, Etosha NP, Namibia, January

Tony Heald / Black rhinoceros {Diceros bicornis} drinking at Okaukuejo waterhole at night, Etosha NP, Namibia, January

Tony Heald / Hippopotamus {Hippopotamus amphibius} mud covered mother and baby, Chobe National Park, Botswana

Tony Heald / African darter {Anhinga rufa} preening wings, Chobe National Park, Botswana

Tony Heald / Hippopotamus {Hippopotamus amphibius} with Red billed oxpeckers on head, Moremi Wildlife Reserve, Botswana

Tony Heald / Hippopotamus {Hippopotamus amphibius}, Moremi Wildlife Reserve, Botswana

Tony Heald / Hippopotamus {Hippopotamus amphibius} running with Oxpecker on its back, Chobe National Park, Botswana

Tony Heald / Hippopotamus {Hippopotamus amphibius} emerging from water, Chobe National Park, Botswana

Tony Heald / Dust storm Kalahari Gemsbok NP South Africa, summer

Tony Heald / Lechwe male head portrait {Kobus leche} Moremi WR Botswana

Tony Heald / Wildebeest fighting (Connochaetes taurinus) Kruger NP, S. Africa

Tony Heald / African buffalo silhouetted at dawn {Syncerus caffer} Moremi NP, Botswana

Tony Heald / African elephant charging (Loxodonta africana) Etosha NP, Namibia

Tony Heald / African elephant {Loxodonta africana} charging abstract, Etosha national park, Namibia. Digitally enhanced version of 01154692.

Tony Heald / African elephant {Loxodonta africana}, close-up of eye, Chobe national park, Botswana

Tony Heald / African elephant {Loxodonta africana} charging, Etosha national park, Namibia.

Tony Heald / Common zebra {Equus quagga} rolling on ground, dustbathing, Etosha national park, Namibia.

Tony Heald / African elephant {Loxodonta africana} kicking up white mud (Calcrete and water) Etosha national park, Namibia.

Tony Heald / Black backed jackal {Canis mesomelas} trying to catch doves, Etosha national park, Namibia.

Tony Heald / White rhinoceros {Ceratotherium simium} head on, Etosha national park, Namibia.

Tony Heald / Black rhinoceros {Diceros bicornis} charging with mouth open, Etosha national park, Namibia

Tony Heald / Female Red footed falcon (western) {Falco vespertinus} in flight, Etosha national park, Namibia.

Tony Heald / African elephant {Loxodonta africana} bulls walking in line, Etosha national park, Namibia.

Tony Heald / Aerial view of sand dune landscape, Namib Naukluft NP, Namibia

Tony Heald / Bat Eared Fox emerging from den (Otocyon megalotis) Kalahari Gemsbok NP South Africa

Tony Heald / Lappet faced vulture {Torgos tracheliotus} being attacked by an aggresive Black backed jackal {Canis mesomelas} Etosha NP, Namibia

Tony Heald / African fish eagle about to catch fish {Haliaeetus vocifer} Chobe NP, Botswana.

Tony Heald / African fish eagle {Haliaeetus vocifer} swooping down in flight, Chobe NP, Botswana

Tony Heald / Circle round the sun, optical effect of high level ice crystals, around midday, Chobe NP, Botswana

Tony Heald / Gabar Goshawk {Micronisus gabar} swooping on prey, Etosha NP, Namibia

Tony Heald / African masked weaver {Ploceus velatus} male in early stage of building nest, Etosha NP, Namibia

Tony Heald / African lion {Panthera leo} female portrait, Laikipia, Kenya

Tony Heald / Reticulated giraffe {Giraffa cameloparalis reticulata} mother and young running, Laikipia, Kenya