Sven Zacek / Brown bear (Ursus arctos) in coniferous forest. Ida-Virumaa, Eastern Estonia. September.
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Nick Upton / Eurasian beaver (Castor fiber) dam made from cut branches, surrounded by wet woodland. Near Lisaku, Ida-Viru County, Estonia. April.
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Nick Upton / Scots Pine tree (Pinus sylvestris) forest on northern shore of Lake Peipsi, near Alajoe, Ida-Viru County, Estonia. April 2018.
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Nick Upton / Wind-blown lake ice piled up near shore, Lake Peipsi, Estonia. April 2018.
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Sven Zacek / Silhouette of Red squirrel (Sciurus vulgaris) gathering nesting material for winter nest. Ida-Virumaa, Eastern Estonia. November.
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Sven Zacek / Brown bear (Ursus arctos) in coniferous forest. Ida-Virumaa, Eastern Estonia. September.
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Sven Zacek / Narva River, the border between Estonia and Russia with medieval castles on each side, in evening light. Ida-Virumaa, Estonia. May 2016.
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Sven Zacek / Pattern of Conifer (Pinaceae) needles on snow, revealed following spring melt. Agusalu Nature Reserve, Ida-Virumaa, Eastern Estonia. April.
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Sven Zacek / Pine (Pinus sp) forest canopy viewed from below. Agusalu Nature Reserve, Ida-Virumaa, Eastern Estonia. April.
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Nick Upton / Felled conifer trees stacked beside forestry track through mixed forest. Near Muraka Forest Reserve, a rare surviving home for Siberian flying squirrel (Pteromys volans). Near Lisaku, Estonia. April 2...
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Nick Upton / Aspen (Populus tremula) with Aspen trunk rot / False tinder conk mushroom (Phellinus tremulae). Trees with rotten cores provide ideal nest holes for the Siberian flying squirrel (Pteromys volans). Mur...
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Nick Upton / Researcher radiotracking Siberian flying squirrel (Pteromys volans) in mature mixed forest. Muraka Forest Reseve, near Lisaku, Estonia. April 2018.
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Nick Upton / Researcher setting trailcam to monitor Siberian flying squirrel (Pteromys volans) nest hole. Mature mixed forest, near Lisaku, Estonia. April 2018. Model released.
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Nick Upton / Researcher using laptop to check trailcam footage of Siberian flying squirrel (Pteromys volans) nest box in mature mixed forest. Near Lisaku, Estonia. April 2018. Model released.
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Nick Upton / Nest box for Siberian flying squirrel (Pteromys volans) in mature mixed forest. Muraka Forest Reseve, near Lisaku, Estonia. April 2018.
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Nick Upton / Mixed wet woodland including Aspen (Populus tremula) and Downy birch (Betula pubescens). Inhabited by rare surviving Siberian flying squirrel (Pteromys volans) population. Near Lisaku, Estonia. April...
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Nick Upton / Mixed wet woodland including Aspen (Populus tremula) and Downy birch (Betula pubescens). Inhabited by rare surviving Siberian flying squirrel (Pteromys volans) population. Near Lisaku, Estonia. April...
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Nick Upton / Siberian flying squirrel (Pteromys volans) caught in emergence trap over nest hole entrance in Aspen (Populus tremula) tree. Muraka Forest Reserve, near Lisaku, Estonia. April.
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Nick Upton / Siberian flying squirrel (Pteromys volans) droppings. Found below an old deciduous tree with nest hole. Muraka Forest Reserve, near Lisaku, Estonia. April.
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Nick Upton / Laptop screen showing trailcam footage of Siberian flying squirrel (Pteromys volans) returning to nest hole in Aspen (Populus tremula) tree. Near Lisaku, Estonia. April 2018.
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Nick Upton / Felled Aspen (Populus tremula) trees with tit nests in hollows. Treeholes suitable for Siberan flying squirrel (Pteromys volans). Near Lisaku, Estonia. April 2018.
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Nick Upton / Downy birch (Betula pubescens) trees in mixed woodland, the tree on the left houses a Siberian flying squirrel (Pteromys volans) nest hole. Near Lisaku, Estonia. April.
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Nick Upton / Researcher radio tracking Siberian flying squirrel (Pteromys volans) in mature mixed forest. Muraka Forest Reseve, near Lisaku, Estonia. April 2018. Model released.
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Nick Upton / Researcher radio tracking radio collared Siberian flying squirrel (Pteromys volans). Mature mixed forest, Muraka Forest Reseve, near Lisaku, Estonia. April 2018. Model released.
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Nick Upton / Felled trees stacked beside forestry track through mixed forest. A few metres outside protected limits of Muraka Forest Reserve, a remaining habitat of Siberian flying squirrel (Pteromys volans). Esto...
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Nick Upton / Siberian flying squirrel (Pteromys volans) caught in trap, prior to fitting of radio collar. Muraka Forest Reserve, near Lisaku, Estonia. April.
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Nick Upton / Researcher labelling collection tube of Siberian flying squirrel (Pteromys volans) droppings. Droppings found below old deciduous tree with suitable nest hole. Muraka Forest Reserve, near Lisaku, Ida...
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Nick Upton / Siberian flying squirrel (Pteromys volans) droppings in moss. Found underneath old deciduous tree with an occupied nest hole. Lisaku Park Forest, Estonia. April.
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Nick Upton / Felled conifers stacked at edge of forest.Near Muraka Forest Reserve, a rare surviving home for the Siberian flying squirrel (Pteromys volans). Estonia. April 2018.
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Nick Upton / Felled Aspen (Populus tremula) trees with tit nests in hollows. Treeholes suitable for Siberan flying squirrel (Pteromys volans). Near Lisaku, Estonia. April 2018.
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Nick Upton / Eurasian aspen (Populus tremula) catkins, major food souce for Siberian flying squirrel (Pteromys volans). Lisaku Park Forest, Estonia. April.
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Nick Upton / Research team preparing to radio collar Siberian flying squirrel (Pteromys volans) caught in a trap. Muraka Forest Reserve, near Lisaku, Estonia. April 2018. Model released.
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Nick Upton / Researcher reviewing trailcam footage on laptop of Siberian flying squirrel (Pteromys volans) emerging from nest hole in Aspen (Populus tremula) tree at night. Mature mixed forest, near Lisaku, Estoni...
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Nick Upton / Researcher examining Siberian flying squirrel (Pteromys volans) droppings. Mature mixed forest, near Lisaku, Estonia. April 2018. Model released.
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Nick Upton / Researcher wearing thick gloves collecting Siberian flying squirrel (Pteromys volans) from trap prior to fitting radio collar. Muraka Forest Reserve, Lisaku, Estonia. April 2018. Model released.
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Nick Upton / Researcher face to face with Siberian flying squirrel (Pteromys volans) whilst setting trap on old Aspen (Populus tremula) tree. Muraka Forest Reserve, near Lisaku, Estonia. April 2018. Model released...
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Nick Upton / Researcher collecting Siberian flying squirrel (Pteromys volans) droppings below an old deciduous tree with an occupied nest hole. Muraka Forest Reserve, near Lisaku, Estonia. April 2018. Model releas...
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Nick Upton / Mixed wet woodland including Aspen (Populus tremula) and Downy birch (Betula pubescens). Inhabited by a surviving population of Siberian flying squirrel (Pteromys volans). Near Lisaku, Estonia. April...
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Nick Upton / Siberian flying squirrel (Pteromys volans) in the hand, prior to fitting of radio collar. Muraka Forest Reserve, near Lisaku, Estonia. April.
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Nick Upton / Siberian flying squirrel (Pteromys volans) entering nest hole in Downy birch (Betula pubescens) tree trunk. Mature mixed forest, near Lisaku, Ida Virumaa, Estonia. April.
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Nick Upton / Researcher examining and sexing Siberian flying squirrel (Pteromys volans) prior to fitting radio collar. Muraka Forest Reserve, near Lisaku, Estonia. April 2018. Model released.
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Nick Upton / Researcher fitting radio collar on Siberian flying squirrel (Pteromys volans) caught in mature mixed forest at night. Near Lisaku, Estonia. April 2018.
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Nick Upton / Siberian flying squirrel (Pteromys volans) peering out of nest hole in old Aspen (Populus tremula) tree. Mature mixed forest, near Lisaku, Estonia. April.
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Nick Upton / Researcher inspecting a Siberian flying squirrel (Pteromys volans) caught in trap on old Aspen (Populus tremula) tree with occupied nest hole. Muraka Forest Reserve, near Lisaku, Estonia. April 2018....
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Sven Zacek / Frozen grassland at sunrise, Agusalu Nature Reserve, Ida-Virumaa, Estonia, November.
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Sven Zacek / Windswept ice and sand patterns on the shore of Lake Peipsi, Ida-Virumaa, Estonia, November.
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Sven Zacek / Aerial view of sandy beach, Kauki on the north coast of Lake Peipsi, Ida-Virumaa, Estonia, November 2014.
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Sven Zacek / Aerial view of sandy beach of Kauksi, Lake Peipsi in Ida-Virumaa, Estonia, October 2014.
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Sven Zacek / Aerial view of bog pools at sunrise, Sirtsi Nature Resrve in Ida-Virumaa, Estonia, May 2014.
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Sven Zacek / Shadows and sunlight in fog over bog at the forest edge, Sirtsi Nature Resrve in Ida-Virumaa, Estonia, May 2014.
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Sven Zacek / Aerial view of bog pools in the mist at sunrise, Sirtsi Nature Reserve, Ida-Virumaa, Estonia, May 2014.
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Sven Zacek / Aerial view of foggy bog pools at sunrise. Ida-Virumaa, Estonia, May.
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Sven Zacek / Snowfall in a pristine boreal forest, Sirtsi Nature Reserve in Ida-Virumaa, Estonia. November 2013.
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Sven Zacek / Aerial view of Muraka bog in Ida-Virumaa county, Estonia, August 2011.
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Sven Zacek / Aerial view of Muraka bog in Ida-Virumaa. Estonia, August 2011.
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Sven Zacek / Clouds moving over frozen Valaste Waterfall, Ida-Virumaa county, Estonia.
