Paul Williams / Tourists watching two wild Jaguar cubs fight on riverbank whilst third looks on from above (Panthera onca) from boat on river, Pantanal, Pocone, Brazil. No release available.

Mark Carwardine / Jaguar (Panthera onca), one-year cub peering from behind tree, Cuiaba River, Pantanal, Brazil. near threatened species

Luiz Claudio Marigo / Female Jaguar (Panthera onca palustris) on the shore of Piquiri River. The Pantanal wetlands of Mato Grosso State, Brazil, October.

Luiz Claudio Marigo / Jaguar (Panthera onca palustris) in water with Yacare Caiman (Caiman yacare), its teeth around the back of the prey's neck. Piquiri River, The Pantanal wetlands of Mato Grosso State, Brazil, October....

Luiz Claudio Marigo / Jaguar (Panthera onca palustris) dragging a Yacare Caiman (Caiman yacare) onto the bank of the Piquiri River. The Pantanal wetlands of Mato Grosso State, Brazil, October.

Luiz Claudio Marigo / Jaguar (Panthera onca palustris) struggling with a Yacare Caiman (Caiman yacare) in the Piquiri River. The Pantanal wetlands of Mato Grosso State, Brazil, October. Sequence 2/4

Luiz Claudio Marigo / Jaguar (Panthera onca palustris) male drinking water from the Piquiri River. The Pantanal wetlands of Mato Grosso State, Brazil, October.

Luiz Claudio Marigo / Jaguar (Panthera onca palustris) male wading through water at the shore of the Piquiri River. The Pantanal wetlands of Mato Grosso, Mato Grosso State, Brazil, October.

Nick Garbutt / Wild female Jaguar (Panthera onca palustris) jumping up sandy bluff on the banks of the Piquiri River (a tributary of Cuiaba River). Northern Pantanal, Brazil. September

Nick Garbutt / Wild female Jaguar (Panthera onca palustris) resting on the banks of the Piquiri River (a tributary of Cuiaba River). Northern Pantanal, Brazil.

Edwin Giesbers / Jaguar (Panthera onca) close-up head portrait, lying down, captive

Edwin Giesbers / Jaguar (Panthera onca) close-up of coat and tail, captive

Edwin Giesbers / Jaguar (Panthera onca) head portrait,rear view, showing ear spots, captive

Edwin Giesbers / Jaguar (Panthera onca) portrait, lying down, captive

Christophe Courteau / Photo-identification of Jaguars (Panthera onca) for scientific Research, Pantanal, Brazil, September 2008

Christophe Courteau / Jaguar (Panthera onca palustris) wild male, on the bank of the Cuiaba River, Pantanal, Mato Grosso, Brazil, September 2008 This individual is called Jack.

Christophe Courteau / Jaguar (Panthera onca palustris) wild pregnant female, in undergrowth beside the Cuiaba River, Pantanal, Mato Grosso, Brazil, September 2008. This individual is called BORBOLETTA by researchers (pho...

Christophe Courteau / Jaguar (Panthera onca palustris) wild pregnant female, in undergrowth beside the Cuiaba River, Pantanal, Mato Grosso, Brazil, September 2008 This individual is called BORBOLETTA by researchers (phot...

Andy Rouse / Jaguar (Panthera onca) on tree with paw in air, in rainforest, captive, Belize

Andy Rouse / Jaguar (Panthera onca) by rainforest pool with reflection in water, captive, Belize

Pete Oxford / Female Jaguar (Panthera onca) under anesthetic, having teeth measured as part of the collection of biometric data and physiological samples for a relocation project, captive, Fundacion Santa Martha Ce...

Edwin Giesbers / Jaguar (Panthera onca) portrait, Costa Rica, Captive

LYNN M. STONE / Jaguar (Panthera onca) Belize captive

Luiz Claudio Marigo / Jaguars fighting {Panthera onca} tropical rainforest, Brazil. Captive.

Hermann Brehm / Jaguar (Panthera onca) male yawning, Pantanal, Brazil

Hermann Brehm / Jaguar (Panthera onca) male walking, Pantanal, Brazil

Hermann Brehm / Jaguar (Panthera onca) male walking near waters edge, Pantanal, Brazil

Hermann Brehm / Jaguar (Panthera onca) male walking and snarling, Pantanal, Brazil

Hermann Brehm / Jaguar (Panthera onca) male standing near river edge, Pantanal, Brazil

Hermann Brehm / Jaguar (Panthera onca) female snarling, Pantanal, Brazil

Hermann Brehm / Jaguar (Panthera onca) female rolling, Pantanal, Brazil

David Pattyn / Jaguar (Panthera onca) resting on a riverbank in Pantanal, Brazil.

David Pattyn / Jaguar (Panthera onca) resting on a riverbank in Pantanal, Brazil.

David Pattyn / Jaguar (Panthera onca) yawning and resting along riverbank in Pantanal, Brazil.

David Pattyn / Jaguar (Panthera onca) resting on a riverbank in Pantanal, Brazil.

David Pattyn / Jaguar (Panthera onca) resting on a sandy river beach in Pantanal, Brazil.

David Pattyn / Jaguar (Panthera onca) resting on a riverbank in Pantanal, Brazil.

David Pattyn / Jaguar (Panthera onca) stalking in high vegetation along a river in search of prey, Pantanal, Brazil.

David Pattyn / Jaguar (Panthera onca) resting on a riverbank in Pantanal, Brazil

Mark Carwardine / Jaguar (Panthera onca) resting, grooming, on riverbank, Cuiaba River, Pantanal, Brazil

Mark Carwardine / Jaguar (Panthera onca) resting on riverbank, Cuiaba River, Pantanal, Brazil *Digitally removed wound on Jaguar's face

Mark Carwardine / Jaguar (Panthera onca) walking along riverbank, Cuiaba River, Pantanal, Brazil *Digitally removed wound on jaguar's face

Mark Carwardine / Jaguar (Panthera onca) walking through vegetation, Cuiaba River, Pantanal, Brazil *Digitally removed wound on jaguar's face

Mark Carwardine / Jaguar (Panthera onca) stretching, Cuiaba River, Pantanal, Brazil *Digitally removed wound on jaguar's face

Mark Carwardine / Jaguar (Panthera onca) looking up, snarling, Cuiaba River, Pantanal, Brazil

Mark Carwardine / Jaguar (Panthera onca) looking up, watching fly, Cuiaba River, Pantanal, Brazil

Mark Carwardine / Jaguar (Panthera onca) looking up, watching fly, Cuiaba River, Pantanal, Brazil

Mark Carwardine / Jaguar (Panthera onca) snarling, Cuiaba River, Pantanal, Brazil *Digitally removed wound on jaguar's face

Mark Carwardine / Jaguar (Panthera onca) Cuiaba River, Pantanal, Brazil *Digitally removed wound on jaguar's face

Mark Carwardine / Jaguar (Panthera onca) licking paw on riverbank, Cuiaba River, Pantanal, Brazil *Digitally removed branch from foreground

Mark Carwardine / Jaguar (Panthera onca) grooming paw on riverbank, wound on face, Cuiaba River, Pantanal, Brazil *Digitally removed branch from foreground

Mark Carwardine / Jaguar (Panthera onca) on riverbank grooming face with paw, Cuiaba River, Pantanal, Brazil *Digitally removed branch from foreground

Mark Carwardine / Jaguar (Panthera onca) amongst vegetation on riverbank, Cuiaba River, Pantanal, Brazil *Digitally removed branch from foreground and wound on jaguar's face

Mark Carwardine / Jaguar (Panthera onca) yawning, Cuiaba River, Pantanal, Brazil *Digitally removed branch from foreground

Mark Carwardine / Jaguar (Panthera onca) walking along riverbank, Cuiaba River, Pantanal, Brazil

Mark Carwardine / Photographers watching a Jaguar (Panthera onca) on riverbank, Cuiaba River, Pantanal, Brazil, July 2009

Mark Carwardine / Jaguar (Panthera onca) one-year cub amongst vegetation near the Cuiaba River, Pantanal, Brazil

Mark Carwardine / Jaguar (Panthera onca) one-year cub amongst vegetation near the Cuiaba River, Pantanal, Brazil

Mark Carwardine / Jaguar (Panthera onca) grooming, resting beside Cuiaba River, Pantanal, Brazil *Digitally removed wound on Jaguar's face

Mark Carwardine / Jaguar (Panthera onca) grooming, resting beside Cuiaba River, Pantanal, Brazil *Digitally removed wound on Jaguar's face

Mark Carwardine / Jaguar (Panthera onca) with wound on face, appearing to snarl, Cuiaba River, Pantanal, Brazil

Mark Carwardine / Jaguar (Panthera onca) resting beside the Cuiaba River, Pantanal, Brazil *Digitally removed wound on Jaguar's face

Mark Carwardine / Jaguar (Panthera onca) sleeping beside the Cuiaba River, Pantanal, Brazil *Digitally removed wound on Jaguar's face

Mark Carwardine / Jaguar (Panthera onca) large male with wound on his face, resting beside the Cuiaba River, Pantanal, Brazil

Mark Carwardine / Jaguar (Panthera onca) resting on river bank beside the Cuiaba River, Pantanal, Brazil *Digitally removed wound on Jaguar's face

Mark Carwardine / Jaguar (Panthera onca) sleeping on river bank beside the Cuiaba River, Pantanal, Brazil

Mark Carwardine / Jaguar (Panthera onca) sleeping beside the Cuiaba River, Pantanal, Brazil *Digitally removed wound on Jaguar's face

Mark Carwardine / Jaguar (Panthera onca) sleeping beside the Cuiaba River, Pantanal, Brazil *Digitally removed wound on Jaguar's face

Mark Carwardine / Jaguar (Panthera onca) resting beside the Cuiaba River, Pantanal, Brazil *Digitally removed wound on Jaguar's face

Paul Williams / Tourists watching wild Jaguar (Panthera onca) from boat on river, Pantanal, Pocone, Brazil

Paul Williams / Jaguar (Panthera onca) yawning whilst at rest on bank, Pantanal, Pocone, Brazil

Paul Williams / Jaguar (Panthera onca) resting on tree trunk, Pantanal, Pocone, Brazil

Paul Williams / Jaguar (Panthera onca) resting on bank, Pantanal, Pocone, Brazil

Paul Williams / Tourists watching two wild Jaguar cubs fight on riverbank, (Panthera onca) from boat on river, Pantanal, Pocone, Brazil

Paul Williams / Tourists watching two wild Jaguar cubs fight on riverbank, (Panthera onca) from boat on river, Pantanal, Pocone, Brazil

Paul Williams / Jaguar (Panthera onca) pug mark next to Apple snail (Pomacea canaliculata) Pantanal, Pocone, Brazil

Paul Williams / Jaguar (Panthera onca) resting in amongst thick vegetation, Pantanal, Pocone, Brazil

Paul Williams / Jaguar (Panthera onca) resting in amongst thick vegetation, Pantanal, Pocone, Brazil

Paul Williams / Jaguar (Panthera onca) pug mark next to Apple snail (Pomacea canaliculata) Pantanal, Pocone, Brazil

Paul Williams / Jaguar (Panthera onca) resting in amongst thick vegetation, Pantanal, Pocone, Brazil

Mark Carwardine / Jaguar (Panthera onca), one-year cub watching a fly from behind tree, Cuiaba River, Pantanal, Brazil. near threatened species

Mark Carwardine / Jaguar (Panthera onca), one-year cub walking through vegetation, Cuiaba River, Pantanal, Brazil. near threatened species

Mark Carwardine / Jaguar (Panthera onca), one-year cub looking down from tree, Cuiaba River, Pantanal, Brazil. near threatened species

Patricio Robles Gil / Female Jaguar (Panthera onca) on river bank. Parana, Southern Brazil.

Patricio Robles Gil / Female Jaguar (Panthera onca) with Jacare Caiman prey. Parana, Southern Brazil.

Patricio Robles Gil / Tourists watching a female Jaguar (Panthera onca) from a boat. Piquiri River, Parana, Southern Brazil.

Patricio Robles Gil / Female Jaguar (Panthera onca) with her prey, a Yacare Caiman. Parana, Southern Brazil.

Patricio Robles Gil / Male Jaguar (Panthera onca) swimming across the River Piquiri wearing a radio location collar. Parana, Southern Brazil.

Jeff Rotman / Red paint covers the top half of a Matses woman's face, and she wears decorative spines on both sides of her nose. This makeup represents the whiskers of the jaguar spirit. Amazon, Peru, November 2005...

Jeff Rotman / Red paint covers the top half of a Matses woman's face, and she wears decorative spines on both sides of her nose. This makeup represents the whiskers of the jaguar spirit. Amazon, Peru, November 200...

Jeff Rotman / Red and black paint covers a Matses woman's face, and she wears decorative spines on both sides of her nose. This makeup represents the whiskers of the jaguar spirit. Amazon, Peru, November 2005.

Jeff Rotman / Red and black paint covers a Matses woman's face, and she wears decorative spines on both sides of her nose. This makeup represents the whiskers of the jaguar spirit. Amazon, Peru, November 2005.

Nature Production / Close up of fur pattern of Jaguar (Panthera onca) captive

Luiz Claudio Marigo / Jaguar (Panthera onca palustris) female on the shore of Piquiri River. The Pantanal wetlands of Mato Grosso State, Brazil, October.