Floris van Breugel / Monarch butterflies (Danaus plexippus) over winter on Eucalyptus and Monterey Cypress trees near Pismo Beach, San Luis, California, USA December.. When temperatures warm up enough, they will embark on...

Olga Kamenskaya / View of people standing on ice from underneath car, on the ice of Lake Baikal, Siberia, Russia, March. Property released and Model released.

Olga Kamenskaya / Mini van crossing Lake Baikal ice, Siberia, Russia, March.

Olga Kamenskaya / Car on thick Lake Baikal ice, Lake Baikal, Siberia, Russia, March. Property released.

Peter Cairns / American bison (Bison bison) walking through snow using tracks made by other animals, Yellowstone National Park, Wyoming, USA, February.

Peter Cairns / American Bison (Bison bison) walking through deep snow, Yellowstone National Park, Wyoming, USA, February.

Adrian Davies / Trail left in a Bramble leaf by the larvae of a Pygmy eyecap moth (Stigmella aurella), scanned on a flatbed scanner, Surrey, England, UK.

Markus Varesvuo / Common shag (Phalacrocorax aristotelis) flying, with the sun in the background, Vardo, Norway, March.

Alex Mustard / Blue shark (Prionace glauca) near the surface of the English Channel. Penzance, Cornwall, England, British Isles. North East Atlantic Ocean, August. Digitally manipulated to improve composition see 1...

Theo Bosboom / Reindeer (Rangifer tarandus) male walking across beach in winter, near Hofn, Iceland, February

Orsolya Haarberg / Chamois (Rupicapra rupicapra) walking, casting shadow on the snow, Alps, Gran Paradiso National Park, Italy. November 2008

Graham Eaton / Leatherback Turtle Hatchling (Dermochelys coriacea) crossing a beach towards the sea at sunrise, Cayenne, French Guiana, July

Claudio Contreras / View from bangka boat at dawn, Danajon Bank, Central Visayas, Philippines, April 2013

Dr. Gertrud Neumann-Denzau / Urial (Ovis orientalis arkal) herd standing on rocks, Touran Protected Area, now part of Khar Turan National Park, Semnan Province, Iran

Michael Pitts / Mike de Gruy, underwater camerman, in Remora submersible, Mediterranean

David Pattyn / Blackbird (Turdus merula) male in flight during snowfall, Oisterwijk, The Netherlands. January

Andy Rouse / Barn Owl (Tyto alba) in flight in dawn light. UK, Europe.

Jason Venus / Western diamondback rattlesnake {Crotalus atrox} in road at night. BBC Incredible Journeys. Road in the path of snake's migratory route across the Sonoran Desert, Arizona USA. Many killed on road.

Jason Venus / Swallow in flight, taken in flight tunnel on loction for BBC television series 'Incredible Journeys'

Jen Guyton / Researcher weighing wild, habituated Meerkats (Suricata suricatta) at their burrow in the Kalahari Desert, South Africa. June 2010. Siena International Photo Awards 2021 Honorable mention - Journeys...

Iris Thorsteinsdottir / Icelandic road in mist, Dettifoss, Iceland, July 2012.

Merryn Thomas / Man in vehicle, photographing Salar de Uyuni salt flats, Atacama Desert, Bolivia. February 2014.

Jordi Chias / Mobula / Chilean devil ray (Mobula tarapacana) with a school of fish (Capros aper) showing vertical migration, Santa Maria, Azores, Portugal, Atlantic Ocean

Eric Baccega / Nenet herder drives Reindeer (Rangifer tarandus) sleds on spring migration across tundra. Yar-Sale district, Yamal, Northwest Siberia, Russia. April 2016.

Eric Baccega / Ekaterina Yaptik, Nenet woman leading Reindeer (Rangifer tarandus) on spring migration in tundra with child in sled. Yar-Sale district, Yamal, Northwest Siberia, Russia. April 2016.

Enrique Lopez-Tapia / Salt caravan of dromedary camels (Camelus dromedarius), crossing Lake Assale, one of the largest salt lakes in Africa. Afar Region, Ethiopia, Africa. November 2014.

Enrique Lopez-Tapia / Man on horseback, with towel and cap on head, Bale Mountains National Park. Ethiopia, November 2014

Cyril Ruoso / Rainbow over road on the south east coast of Iceland, August 2003.

Ole Jorgen Liodden / Polar bear (Ursus maritimus) walking on ice, reflected in still water, Spitsbergen, Svalbard, Norway. July.

Ole Jorgen Liodden / Prow of expedition ship with water splashing around it, Drake Passage, Chile. March 2006.

Markus Varesvuo / Whooper Swan (Cygnus cygnus) flock in flight over mountains, Hokkaido, Japan, February

Frederique Olivier / Adelie penguin (Pygoscelis adeliae) group crossing ice to sea edge to feed, Antarctica.

Frederique Olivier / Adelie penguins (Pygoscelis adeliae) travelling to the ice edge to feed, Antarctica.

Frederique Olivier / Adelie penguin (Pygoscelis adeliae) group travelling to edge of ice to feed, Antarctica.

Chris Mattison / US Highway 198 passing through the Sequoia National Park, California, USA, May.

David Noton / Avenue of Plane trees (Platanus) on a road near Soreze, Tarn, Languedoc, France, September 2012.

David Noton / Avenue of Plane trees (Platanus) on a road near Soreze, Tarn, Languedoc, France, September 2012.

David Noton / Avenue of Plane trees (Platanus) on a road near Soreze, Tarn, Languedoc, France, September 2012.

David Noton / Avenue of Plane trees (Platanus) on a road near Soreze, Tarn, Languedoc, France, September 2012.

David Noton / Children cycling to school, near Vang Vieng, Laos, March 2009.

David Noton / People crossing a footbridge over the Nam Khan River, Luang Prabang, Laos, March 2009.

David Noton / Children riding bikes along riverside path at dawn, near Vang Vieng, Laos, March 2009.

David Noton / Cyclists riding bikes with tractor along riverside path at dawn, near Vang Vieng, Laos, March 2009.

David Noton / Road to Keno, with snow capped mountain and Quaking aspen (Populus tremuloides) Silver Trail near Mayo, Yukon Territories, Canada, September 2013.

David Noton / The road to Skagway with autumnal trees and mountain, South Klondike Highway, Yukon Territories, Canada, September 2013.

David Noton / The road to Skagway with autumnal trees and mountain, South Klondike Highway, Yukon Territories, Canada, September 2013.

Steve O. Taylor (GHF) / West African refugees crossing the Sahara on their way to Europe, Niger, 2005.

Steve O. Taylor (GHF) / Man weaving traditional Peul / Fula fabric on loom. National Museum, Naiamey, Niger, 2004.

Steve O. Taylor (GHF) / Peul / Fula woman travelling to new encampment on donkey, central Niger, 2004.

Steve O. Taylor (GHF) / Refugees from West Africa crossing the Sahara on their way to Europe, Niger, 2005.

Steve O. Taylor (GHF) / West African refugees travelling north to Libya across the Sahara and then onto Europe. Niger, 2005.

Graham Eaton / Leatherback Turtle Hatchling (Dermochelys coriacea) crossing a beach towards the sea at dawn, Cayenne, French Guiana, July

Graham Eaton / Leatherback Turtle Hatchling (Dermochelys coriacea) crossing a beach towards the sea, Cayenne, French Guiana, July

Pascal Tordeux / A path from the peaks in Vars Queyras massif, above the park Escreins, France, August 2011

Paul Harcourt Davies / Ancient Roman road made in Basalt near Bolsena, Lazio. Italy, January

Juergen Freund / Birdsville Track, South Australia, Australia

Juergen Freund / Roadtrains traversing the Birdsville Track, Queensland, Australia, June 2011

George Sanker / Appalachain Trail running through forest in Baxter State Park, Maine, USA. October 2012.

Kirkendall-Spring / Road #653 in the Sawtooth National Recreation Area. Idaho, USA, July 2011

Bryan and Cherry Alexander / Chukchi hunters travelling from Uelen to Dezhnovka by dog sled on a windy November day. Uelen, Chukotka, Siberia, 2006. 40 BELOW bookplate.

Jeff Vanuga / Road stretching towards dramatic mountain peaks. Grand Teton National Park, Wyoming, September 2010.

Bryan and Cherry Alexander / Aerial view of Emperor penguins (Aptenodytes forsteri) travelling across snow, Antarctica.

Jason Venus / Filming a swallow in a flight tunnel on location in Morocco, for BBC television series "Incredible Journeys", 1990s

David Noton / Moon high above disappearing road into Monument Valley Tribal Reserve, Utah, USA

Solvin Zankl / Rockhopper penguin {Eudyptes chrysocome} following path up steep section of rock, towards breeding colony. Note distinctive claw marks scratched onto the rocks as a result of millions of journeys made...

Daniel Gomez / Ticket sellers for boat journeys on the Amazonas river, Brazil.

Jason Venus / Geoffrey Bell prepares remote control filming helicopter on location for BBC series Incredible Journeys.

Gavin Hellier / Road leading to Monument Valley, Navajo Tribal Park, Arizona, USA

Paul D Stewart / Portrait of Carl Linnaeus mezzotint engraving by H. Kingsbury with later hand colouring, published by 'Dr. Thornton 1805 from an original in his possesion'. The original was by Martin Hoffman, painted...