Peter Blackwell / Leopard tortoise portrait {Geochelone pardalis} Amboseli NP, Kenya, East Africa

Christophe Courteau / Leopard tortoise (Geochelone pardalis), Okavango Delta, Botswana

Christophe Courteau / Leopard tortoise (Geochelone pardalis) walking on short grass, Okavango Delta, Botswana

Christophe Courteau / African Lioness (Panthera leo) with a Leopard tortoise (Geochelone pardalis) in her mouth, Botswana

Roy Mangersnes / Leopard tortoise (Geochelone pardalis), Okavango Delta, Botswana, November

Roy Mangersnes / Leopard tortoise (Geochelone pardalis), Okavango Delta, Botswana, November

Roy Mangersnes / Leopard tortoise (Geochelone pardalis), Okavango Delta, Botswana, November

Konrad Wothe / Leopard Tortoise (Geochelone pardalis) grazing, Africa

Tony Phelps / Leopard tortoise (Stigmochelys pardalis) female on track in succulent karoo habitat. Oudtshoorn, Little Karoo, Western Cape, South Africa.

Tony Phelps / Leopard tortoise (Stigmochelys pardalis) female, dorsal view of carapace. Oudtshoorn, Little Karoo, Western Cape, South Africa.

Tony Phelps / Leopard tortoise (Stigmochelys pardalis) adult female, ventral view of plastron. Oudtshoorn, Little Karoo, Western Cape, South Africa.

Tony Phelps / Leopard tortoise (Stigmochelys pardalis) pair mating, with another male next to them, deHoop Nature Reserve, Western Cape, South Africa

Tony Phelps / Leopard tortoise (Stigmochelys pardalis) pair mating, with another male next to them, deHoop Nature Reserve, Western Cape, South Africa

Tony Phelps / Leopard tortoise (Stigmochelys pardalis) two males following female, deHoop Nature reserve, Western Cape, South Africa

Ole Jorgen Liodden / Leopard Tortoise (Geochelone pardalis). Okavango, Botswana, November.

Lisa Hoffner / Leopard tortoise (Geochelone pardalis) seeks the only shade it can find, its shell, in the heat of the day on the dry riverbed of the Ewaso Nyiro. Kenya. The worst drought (2008-2009) in more than a...

Richard Du Toit / Leopard Tortoise (Geochelone pardalis) baby walking on road, Northern Cape, South Africa

Tui De Roy / Leopard Tortoise (Geochelone pardalis), Mpala Research Centre, Kenya

Tui De Roy / Leopard Tortoise (Geochelone pardalis), Mpala Research Centre, Kenya

Christophe Courteau / Leopard tortoise (Geochelone pardalis) swimming through flood water, Okavango Delta, Botswana, March.

Anup Shah / Leopard tortoise (Geochelone pardalis) walking through a shallow pool of water. Masai Mara National Reserve, Kenya. March

Anup Shah / Leopard tortoise (Geochelone pardalis) walking through a shallow pool of water. Masai Mara National Reserve, Kenya. March

Edwin Giesbers / Leopard Tortoise {Geochelone pardalis} Tanzania

Tony Phelps / Leopard tortoise (Stigmochelys / Geochelone pardalis) with damged shell, de Hoop Nature reserve, western Cape, South Africa

Tony Phelps / Mountain / Leopard tortoise (Geochelone / Stigmachelys pardalis) Male amongst vegetation. Little Karoo, South Africa.

Tony Heald / Leopard Tortoise (Geochelone pardalis) portrait, Etosha National Park, Namibia, January

Edwin Giesbers / Leopard tortoise (Geochelone pardalis) walking across sand, Tanzania

Tony Phelps / Leopard tortoise (Geochelone / Stigmachelyus pardalis) large male resting in shade, Little Karoo, South Africa

Tony Phelps / Leopard tortoise (Stigmachelys pardalis) adult male, Little Karoo, South Africa

Tony Phelps / Leopard tortoise (Stigmachelys pardalis) adult, Little Karoo, South Africa

Tony Phelps / Leopard tortoise (Geochelone / Stigmachelys pardalis) adult portrait, Little Karoo, South Africa

Jouan & Rius / Leopard tortoise (Geochelone pardalis) Erongo mountains, Namibia

Vincent Grafhorst / Leopard Tortoise (Geochelone pardalis), Botsalano Game Reserve, South Africa

Vincent Grafhorst / Leopard Tortoise (Geochelone pardalis), Botsalano Game Reserve, South Africa

Vincent Grafhorst / Leopard Tortoise (Geochelone pardalis), Botsalano Game Reserve, South Africa

Nick Garbutt / Various skulls of animals (monkey, gorilla, chimpanzee, buffalo, leopard, crocodile, tortoise) killed / poached illegally for the bush meat trade. Loango National Park, Gabon, Central Africa. February...

Nick Garbutt / Various skulls of animals (monkey, gorilla, chimpanzee, buffalo, leopard, crocodile, tortoise) killed / poached illegally for the bush meat trade. Loango National Park, Gabon, Central Africa. February...

Suzi Eszterhas / 3month-old Cheetah cub {Acinonyx jubatus} watching Leopard tortoise {Geochelone pardalis} with interest , Ngorongoro Conservation Area, Tanzania

Suzi Eszterhas / 3month-old Cheetah cub {Acinonyx jubatus} watching Leopard tortoise {Geochelone pardalis} with interest , Ngorongoro Conservation Area, Tanzania

Anup Shah / Leopard tortoise {Geochelone pardalis} Serengeti NP, Tanzania

Richard Du Toit / Leopard Tortoise (Geochelone pardalis) male at left, female at right, Kgalagadi Transfrontier Park, South Africa

Mike Wilkes / Leopard tortoise {Geochelone pardalis} Kenya

Pete Oxford / Lions {Panthera leo} play with Leopard tortoise, Okavango delta, Botswana

Richard Du Toit / Leopard tortoise drinking {Geochelone pardalis} Kalahari Gemsbok NP, South Africa

Sharon Heald / African Lion {Panthera leo} young male trying to eat Leopard tortoise {Stigmochelys / Geochelone pardalis} Etosha national park, Namibia.

Jane Burton / Leopard tortoise (Geochelone pardalis) running (digitally enhanced)

Tony Phelps / Leopard tortoise (Stigmochelys / Geochelone pardalis) laying egg in nest hole, Oudtshoorn, Little Karoo, South Africa

Tony Phelps / Leopard tortoise {Stigmochelys / Geochelone pardalis} filling in nest hole, Oudtshoorn, Little Karoo, South Africa

Tony Phelps / Leopard tortoise (Stigmochelys / Geochelone pardalis) laying eggs in nest hole, Oudtshoorn, Little Karoo, South Africa

Tony Phelps / Leopard tortoise (Stigmochelys / Geochelone pardalis) laying egg in nest hole, Oudtshoorn, Little Karoo, South Africa

Anup Shah / Leopard juveniles playing with tortoise (Panthera pardus) Kenya, Masai Mara

Andrew Parkinson / Leopard tortoise crossing sandy track {Geochelone pardalis} Chobe, Botswana

Mike Wilkes / Leopard tortoises mating {Geochelone pardalis} Kenya.

Richard Du Toit / Leopard tortoises walking {Geochelone pardalis} Kalahari Gemsbok NP, S Africa

Winfried Wisniewski / Leopard Tortoise (Geochelone pardalis) walking, South Africa

Winfried Wisniewski / Leopard Tortoise (Geochelone pardalis) walking, South Africa

Gerry Ellis / Leopard Tortoise (Geochelone pardalis) on paved road, South Africa

Gerry Ellis / Leopard Tortoise (Geochelone pardalis) portrait, Karoo National Park, west South Africa

Gerry Ellis / Leopard Tortoise (Geochelone pardalis) backside, Karoo National Park, South Africa

Andrew Parkinson / Leopard tortoise at water {Geochelone pardalis} Addo Elephant NP, S Africa

Anup Shah / Leopard cubs playing with tortoise {Panthera pardus} Masai Mara, Kenya

Anup Shah / Young Leopard cub (Manga) trying to feed on tortoise {Panthera pardus} Masai Mara, Kenya

Bernard Castelein / Leopard tortoise on sand. Tsavo National Park, Kenya

Richard Du Toit / Leopard tortoise {Geochelone pardalis} male pursuing female Kalahari Gemsbok NP S Africa. Female the larger.

Richard Du Toit / Leopard tortoise with coral aloes (Aloe striata) flowering. South Africa, Eastern Cape.

Richard Du Toit / Leopard tortoise (Geochelone pardalis) drinking. Kalahari Gemsbok NP, South Africa

Peter Blackwell / Leopard tortoise portrait {Geochelone pardalis} Amboseli NP, Kenya, East Africa

Anup Shah / Leopard tortoise {Geochelone pardalis} pair mating, Kenya, East Africa

Anup Shah / Mating Leopard tortoises {Geochelone pardalis} Kenya, East Africa

Matthias Breiter / Leopard Tortoise (Geochelone pardalis), Linyanti River, Botswana

TJ Rich / Leopard Tortoise (Geochelone pardalis) on savannah, Masai Mara Reserve, Kenya

Andrew Parkinson / Leopard tortoise drinking {Stigmochelys / Geochelone pardalis} Addo Elephant NP S Africa