Nick Upton / Great capricorn / Great oak longhorn beetle (Cerambyx cerdo), one of the largest beetles in Europe, on tree trunk, Lesbos (Lesvos), Greece, June.

Nick Upton / Shoal of young sardines (Sardina pilchardus) swimming in various directions close to the shore in a shallow sandy bay, viewed from above water. Eastern Lesbos /Lesvos, Greece, August.

Nick Upton / Shoaling young sardines (Sardina pilchardus) swimming in shallow water in a sandy bay, casting shadows on the seafloor, viewed from above water. Eastern Lesbos / Lesvos, Greece, August.

Edwin Giesbers / Common tern (Sterna hirundo) flying through grassland at dusk, Lesbos, Greece

Edwin Giesbers / Black-winged stilt (Himantopus himantopus) in flight over long grasses at dusk, Lesbos, Greece

Edwin Giesbers / Small flock of Greater flamingo (Phoenicopterus roseus) wading on inland Kalloni lake, Lesbos, Greece

Edwin Giesbers / Silhouette of Black-winged stilt (Himantopus himantopus) standing on exposed rock over water, with reflections, Lesbos, Greece

Edwin Giesbers / Grecian streamertail / Threadwinged antlion (Nemoptera coa) on flower, Lesbos, Greece

Edwin Giesbers / Portrait of Small skipper butterfly (Thymelicus sylvestris) at rest on flower, Lesbos, Greece

Edwin Giesbers / European pond turtle (Emys orbicularis), Lesbos, Greece

Edwin Giesbers / European pond turtle (Emys orbicularis), swimming with head above water, Lesbos, Greece

David Kjaer / Female Red footed falcon {Falco vespertinus} Lesbos, Greece

Mike Wilkes / Caspian terrapin with young {Mauremys caspica} Lesbos, Greec

George McCarthy / Spur thighed tortoise portrait {Testudo graeca} Island of Lesbos, Greece

Ashley Cooper / Landfill site in Eresos, Lesbos, Greece. June 2011

Ashley Cooper / Landfill site in Eresos, Lesbos, Greece. June 2011

Ashley Cooper / Landfill site in Eresos, Lesbos, Greece. June 2011

Ashley Cooper / Life jackets and remains of boats left by Syrian migrants, refugees fleeing the war. Lesvos Island, Efthalou, Greece. September.

Ashley Cooper / Syrian migrants, refugees fleeing the war, Lesvos Island, Greece. September 2015.

Ashley Cooper / Life jackets and remains of boats left by Syrian migrants, refugees fleeing the war. Lesvos Island, Efthalou, Greece. September.

Ashley Cooper / Solar thermal panels for heating water in Skala Eresou on Lesvos, Greece. June 2013

Ashley Cooper / A swimming pool at a holiday complex in Skala Eresou on Lesvos, Greece. June 2013

Ashley Cooper / Striped necked terrapins (Mauremys caspica) in a river in Skala Eresou, Lesvos, Greece. June.

Ashley Cooper / Beech marten, (Martes foina) killed on a road on Lesvos, Greece. June.

Ashley Cooper / Beech marten, (Martes foina) killed on a road on Lesvos, Greece. June.

Ashley Cooper / An old windmill converted into a house in Sigri on Lesvos, with a fossilised tree in the foreground. Greece, June 2013

Ashley Cooper / Wind farm above Eressos on Lesvos, Greece. June 2013

Hans Glader/ BIA / Kentish Plover (Charadrius alexandrinus) male, Lesvos, Greece

Hans Glader/ BIA / Collared Pratincole (Glareola pratincola), Lesvos, Greece

Hans Glader/ BIA / Collared Pratincole (Glareola pratincola), Lesvos, Greece

Hans Glader/ BIA / Kentish Plover (Charadrius alexandrinus) male, Lesvos, Greece

Hans Glader/ BIA / Little Crake (Porzana parva) male foraging, Lesvos, Greece

Hans Glader/ BIA / Glossy Ibis (Plegadis falcinellus) flying, Lesvos, Greece

Hans Glader/ BIA / Marsh Sandpiper (Tringa stagnatilis) wading, Lesvos, Greece

Mathias Schaef/ BIA / European Goldfinch (Carduelis carduelis), Lesvos, Greece

Mathias Schaef/ BIA / Eastern Orphean Warbler (Sylvia crassirostris) male calling, Lesvos, Greece

Mathias Schaef/ BIA / Woodchat Shrike (Lanius senator) male, Lesvos, Greece

Mathias Schaef/ BIA / Red-rumped Swallow (Cecropis daurica), Lesvos, Greece

Mathias Schaef/ BIA / Red-rumped Swallow (Cecropis daurica), Lesvos, Greece

Mathias Schaef/ BIA / Red-rumped Swallow (Cecropis daurica) flying, Lesvos, Greece

Mathias Schaef/ BIA / Barn Swallow (Hirundo rustica) flying, Lesvos, Greece

Mathias Schaef/ BIA / Isabelline Wheatear (Oenanthe isabellina), Lesvos, Greece

Nick Upton / White-spotted rose beetles (Oxythyrea funesta) mating on Cretan oregano flowers (Origanum onites), Lesbos / Lesvos, Greece, May.

Nick Upton / Turkish meadow brown butterfly (Maniola megala) feeding on Cretan oregano (Origanum onites) flowers with wings closed, Lesbos (Lesvos), Greece, May.

Nick Upton / Tumbling flower beetle / Pintail beetle (Mordellidae) feeding on Cretan oregano flower (Origanum onites) Lesbos / Lesvos, Greece, May.

Nick Upton / Large thistle weevil / Sluggish weevil (Cleonis pigra), on fabric, Lesbos / Lesvos, Greece, May.

Nick Upton / Thick-headed fly (Physocephala rufipes), a parasite of bees, feeding on Japanese mock orange flowers (Pittosporum tobira), Lesbos/ Lesvos, Greece, May.

Nick Upton / Sweat bee (Halictus cf. brunnescens) female foraging on Cretan organo (Origanum onites), Lesbos/ Lesvos, Greece.

Nick Upton / Spurge hawkmoth caterpillar (Hyles euphorbiae) on Broad-leaved glaucous spurge / Myrtle spurge (Euphorbia myrsinites) stripped of its leaves, on limestone mountain top, Mount Olympus, Lesbos / Lesvos,...

Nick Upton / Spurge hawkmoth caterpillar (Hyles euphorbiae) feeding on Sea spurge (Euphorbia paralias) among sand dunes, Lesbos / Lesvos, Greece, May.

Nick Upton / Thread-winged lacewing / Spoonwing lacewing (Nemoptera sinuata) feeding on Cretan oregano flowers (Origanum onites), Lesbos / Lesvos, Greece, May.

Nick Upton / Female Greek paper wasps (Polistes hellenicus) tending grubs in their nest in coastal scrubland, Lesbos / Lesvos, Greece, May.

Nick Upton / Femae Greek paper wasps (Polistes hellenicus) on their nest in coastal scrubland, Lesbos / Lesvos, Greece, May.

Nick Upton / Female Napoleon crab spider (Synema globosum) waiting for prey on Cretan oregano flowerhead (Origanum onites), Lesbos / Lesvos, Greece, May.

Nick Upton / Mediterranean flatheaded woodborer / Jewel beetle (Capnodis cariosa / tenebrionis), a pest of fruit trees including Apricot and Almond, well camouflaged on a tree trunk, Lesbos / Lesvos, Greece, May

Nick Upton / Mediterranean flatheaded woodborer / Jewel beetle (Capnodis cariosa / tenebrionis), a pest of fruit trees including Apricot and Almond, well camouflaged on a tree trunk, Lesbos / Lesvos, Greece, May

Nick Upton / Mediterranean flatheaded woodborer / Jewel beetle (Capnodis cariosa / tenebrionis), a pest of fruit trees including Apricot and Almond, well camouflaged on a tree trunk, Lesbos / Lesvos, Greece, May

Nick Upton / Mammoth wasp / Giant solitary wasp (Megascolia maculata maculata) female sunning on a tree trunk, Lesbos/ Lesvos, Greece, May.

Nick Upton / Lynx spider (Oxyopes heteropthalmus) hunting for insect prey on Cretan oregano flowers (Origanum onites), Lesbos / Lesvos, Greece, May.

Nick Upton / Long-legged bush cricket nymph (Acrometopa syriaca or servillea) standing on a thistle, Mount Olympus, Lesbos / Lesvos, Greece, May.

Nick Upton / Long-legged bush cricket nymph (Acrometopa syriaca or servillea) standing on a Broad-leaved Glaucous Spurge / Myrtle spurge (Euphorbia myrsinites) flower, Mount Olympus, Lesbos / Lesvos, Greece, May.

Nick Upton / Long-legged bush cricket nymph (Acrometopa syriaca or servillea) standing on a Broad-leaved Glaucous Spurge / Myrtle spurge (Euphorbia myrsinites) flower, Mount Olympus, Lesbos / Lesvos, Greece, May.

Nick Upton / Male Ladybird spider (Eresus cinnaberinus / niger) searching for females, Lesbos / Lesvos, Greece, May.

Nick Upton / Male Ladybird spider (Eresus cinnaberinus / niger) searching for females on a rock face, Lesbos / Lesvos, Greece, May.

Nick Upton / Horse fly (Pangonius funebris), a carpenter bee mimic, using its long proboscis to forage on Milk thistle (Carduus marianus), Lesbos / Lesvos, Greece, May.

Nick Upton / Hornet hoverfly / Banded hoverfly (Volucella zonaria) feeding on Japanese mock orange flowers (Pittosporum tobira), Lesbos/ Lesvos, Greece, May.

Nick Upton / Hornet hoverfly / Banded hoverfly (Volucella zonaria) feeding on Japanese mock orange flowers (Pittosporum tobira), Lesbos/ Lesvos, Greece, May.

Nick Upton / Heather crab spider (Thomisus onustus) waiting for insect prey on Cretan oregano flowerhead (Origanum onites), Lesbos/ Lesvos, Greece, May.

Nick Upton / Heather crab spider (Thomisus onustus) waiting for insect prey on Cretan oregano flowerhead (Origanum onites), Lesbos/ Lesvos, Greece, May.

Nick Upton / Female Harvestman (Zacheus crista) well camouflaged on a tree trunk, Lesbos / Lesvos, Greece, May.

Nick Upton / Female Harvestman (Zacheus crista) on a house wall, Lesbos / Lesvos, Greece, May.

Nick Upton / Digger wasp (Sphex pruinosus) male sunning on a rock in hot sunshine and raising its legs, Lesbos/ Lesvos, Greece, May

Nick Upton / Digger wasp (Sphex pruinosus) male sunning on a rock in hot sunshine and raising its legs, Lesbos/ Lesvos, Greece, May

Nick Upton / Copper Buprestid / Jewel beetle (Aurigena / Perotis lugubris) a pest of fruit tee roots, Lesbos / Lesvos, Greece, May

Nick Upton / Copper Buprestid / Jewel beetle (Aurigena / Perotis lugubris) a pest of fruit tee roots, Lesbos / Lesvos, Greece, May

Nick Upton / Young Western Caspian pond turtle / Balkan stripe-necked terrapin (Mauremys caspica rivulata) swimming in a pond with its head above water, Isle of Lesbos / Lesvos, Greece, May.

Nick Upton / Two Western Caspian pond turtles / Balkan stripe-necked terrapins (Mauremys caspica rivulata) swimming in a pond with their heads above water, Isle of Lesbos / Lesvos, Greece, May.

Nick Upton / Bumblee scarab beetle (Eulasia vittata) foraging on a flower, Lesbos / Lesvos, Greece, May

Nick Upton / Bumblee scarab beetle (Eulasia vittata) foraging on a flower, Lesbos / Lesvos, Greece, May

Nick Upton / Bee fly (Thyridanthrax perspicillaris) with strongly patterned wings, feeding on Cretan oregano flowers (Origanum onites), Lesbos / Lesvos, Greece, May.

Nick Upton / Beautiful Jumping spider / Beautiful jumper (Philaeus / Phylaeus chrysops) hunting on rocks behind a beach, Lesbos / Lesvos, Greece, May.

Nick Upton / Beautiful Jumping spider / Beautiful jumper (Philaeus / Phylaeus chrysops) hunting on rocks behind a beach, Lesbos / Lesvos, Greece, May.

Nick Upton / Beautiful Jumping spider / Beautiful jumper (Philaeus / Phylaeus chrysops) hunting among the tide wrack of Seagrass leaves on a beach, Lesbos / Lesvos, Greece, May.

Nick Upton / Lesbian / Balkan marbled white (Melanarge larissa lesbina) feeding from Milk thistle (Silybum marianum = Carduus marianum) near the coast, Lesbos / Lesvos, Greece, May.

Nick Upton / Lesbian / Balkan marbled white (Melanarge larissa lesbina) feeding from Milk thistle (Silybum marianum = Carduus marianum) alongside a Burnet moth (Zygaena punctum), Lesbos / Lesvos, Greece, May.

Nick Upton / Moored fishing boats in Apothika village harbour, with clouds over Mount Olympus in the background, Lesbos/Lesvos, Greece, May 2013.

Nick Upton / Broad-leaved Glaucous Spurge / Myrtle spurge (Euphorbia myrsinites) flowering on limestone mountain top, Mount Olympus, Lesbos, Greece, May 2013.

Nick Upton / Red trefoil / Noble clover (Trifolium rubens) flowering wild in roadside scrub, Lesbos, Greece, May 2013.

Nick Upton / Broad-leaved Glaucous Spurge / Myrtle spurge (Euphorbia myrsinites) flowering on limestone mountain top, Mount Olympus, Lesbos, Greece, May 2013.

Nick Upton / Thermal river, fed with boiling water from hot springs, with colourful growths and scummy crusts of blue-green algae, Lisvori, Polychnitos, Lesbos / Lesvos, Greece, May 2013.

Nick Upton / Thermal river, fed with boiling water from hot springs, with colourful growths and scummy crusts of blue-green algae, Lisvori, Polychnitos, Lesbos / Lesvos, Greece, May 2013.

Nick Upton / Thermal river, fed with boiling water from hot springs, with colourful growths and scummy crusts of blue-green algae, Lisvori, Polychnitos, Lesbos / Lesvos, Greece, May 2013.

Nick Upton / Broad-leaved Glaucous Spurge / Myrtle spurge (Euphorbia myrsinites) clumps flowering on limestone mountain top, Mount Olympus, Lesbos, Greece, May 2013.

Nick Upton / Red trefoil / Noble clover (Trifolium rubens) flowering wild in roadside scrub, Lesbos, Greece, May 2013.

Robin Chittenden / Pelinaio orchid (Ophrys pelinaea) Lesvos, Lesbos, Greece, April.

Robin Chittenden / Pontic fritillary (Fritillaria pontica) Lesbos, Lesvos, Greece, April.