Andy Rouse / Maasai Giraffe (Giraffa camelopardalis tippelskirchi) silhouetted by beams of light from setting sun, Maasai Mara, Kenya, Africa. Winner of the African section of the Natures Best Windland Smith Rice...

Doug Perrine / Bioluminescence from glowing plankton in sea tide line on beach, with stars above and ship lights on horizon, Vaadhoo Island, Raa Atoll, Maldives, Indian Ocean, October 2010

Bence Mate / European rabbits (Oryctolagus cuniculus) fighting each other, Kiskunsag National Park, Hungary. June. Winner, 2019 Cadiz Photo Nature Competition.

Juergen & Christine Sohns / Meerkat (Suricata suricatta) on alert, Tswalu Game Reserve, South Africa

Danny Green / Polar bear (Ursus maritimus) two walking in snowy landscape with glacier behind. Svalbard, Norway, April 2018.

Ann & Steve Toon / Cheetah (Acinonyx jubatus) walking, silhouetted at dusk. Zimanga private game reserve, KwaZulu-Natal, South Africa.

Ann & Steve Toon / African elephants (Loxodonta africana) herd silhouetted at sunset. Chobe National Park, Botswana.

Ann & Steve Toon / Ostrich (Struthio camelus) silhouetted in dust storm. Kalahari Gemsbok National Park, Kgalagadi Transfrontier National Park, Northern Cape, South Africa. Commended in the Bird Photographer of the Year...

Guy Edwardes / Barn owl (Tyto alba) in flight above post and wildflowers, in morning light. The Hawk Conservancy Trust, Hampshire, England, UK. July. Captive.

Guy Edwardes / Glastonbury Tor at sunrise surrounded by early morning mist. View from Walton Hill, Somerset, England, UK. June 2019.

Fabio Pupin / Moorish gecko, (Tarentola mauritanica), hunting on window, Italy, April . Non-ex.

Doug Gimesy / Little blue penguin (Eudyptula minor), two standing on rocks at night, silhouetted against Melbourne city lights. St Kilda breakwater, Victoria, Australia. December 2016. COP26 Countdown Photo Competi...

Heather Angel / Man using pole to propel bamboo raft on Lijang / Li River, Karst peaks beyond. At dawn, Lijiang River Scenic Zone, Yangshuo county, Guangxi, China.

Oscar Dewhurst / Fallow deer (Dama dama) walking across frost-covered playing fields at dawn, with tower blocks of London in the background. Richmond Park, London, UK. December

Oscar Dewhurst / Red deer (Cervus elaphus) stag roaring, silhouetted under trees at sunrise. Bushy Park, London, UK. September

David Woodfall / Sunrise over offshore wind turbine farm, Essex, England, UK, December.

Alex Mustard / Orange painted frogfish (Antennarius pictus) fishing. Here the frogfish is illuminated by a snooted strobe and the background sand lit with a red LED torch. Bitung, North Sulawesi, Indonesia. Lembeh S...

Ernie Janes / Beech trees (Fagus sylvatica) at night in winter, with lamp post illuminating the woodland. Felbrigg Woods, Norfolk, England, UK. January.

Jack Dykinga / Saguaro cacti (Carnegia gigantea) at sunset, during a summer rain storm. Saguaro National Park, Sonoran Desert, Arizona, USA, August. Digital composite.

Espen Bergersen / Killer whales / orcas (Orcinus orca). Spyhopping. Kvaloya, Troms, Norway October

Espen Bergersen / Killer whales / orcas (Orcinus orca). Spyhopping. Kvaloya, Troms, Norway October

Francisco Marquez / Greater flamingo (Phoenicopterus ruber) juveniles and a few adults kept temporarily in an enclosure for ringing, Fuente de Piedra lagoon, Malaga, Spain. August.

SCOTLAND: The Big Picture / Avielochan at sunrise, Cairngorms National Park, Scotland, UK, August.

SCOTLAND: The Big Picture / Red squirrel, (Sciurus vulgaris), silhouetted on stump at dawn, Scotland, UK.March

SCOTLAND: The Big Picture / Red deer, (Cervus elaphus), stag silhouetted at sunset, Scotland, UK.February

SCOTLAND: The Big Picture / Red deer, (Cervus elaphus), stags silhouetted at sunset in winter, Scotland, UK.February

SCOTLAND: The Big Picture / Red deer, (Cervus elaphus), stags silhouetted at sunset in winter, Scotland, UK.February

SCOTLAND: The Big Picture / Red squirrel, (Sciurus vulgaris), two backlit on pine branch, Cairngorms National Park, Scotland, UK.May

SCOTLAND: The Big Picture / Red squirrel, (Sciurus vulgaris), three animals backlit on pine branch, Cairngorms National Park, Scotland, UK.May

SCOTLAND: The Big Picture / Puffin (Fratercula arctica) silhouetted at sunset on Hermaness, Shetland, Scotland, UK, June.

SCOTLAND: The Big Picture / Close up of a colony of Jewel anemones (Corynactis viridis) Scotland, UK, October.

Steven David Miller / Silhouette of an Agile wallaby (Macropus agilis) hopping along the beach at dusk, Cape Hillsborough National Park, Queensland, Australia. September.

Steven David Miller / An Orange lacewing butterfly (Cethosia penthesilea), Cairns Botanical Gardens, Queensland, Australia. Captive. August.

Ross Hoddinott / Banded demoiselles (Calopteryx splendens) male, roosting among grasses, silhouetted against the sun reflected in the water, Lower Tamar Lakes, Cornwall, UK. July.

Ross Hoddinott / Close-up of backlit Porcelain fungus (Oudemansiella mucida) showing gills, Golith Falls, Bodin, Cornwall, UK. September.

Marie Read / Abstract patterns of reflected sky and clouds on the ocean with a silhouetted Red-faced cormorant (Phalarocorax urile) in flight, St. Paul Island, Pribilof Islands, Alaska, USA, July.

Ralph Pace / Green Sea Turtle (Chelonia mydas), San Diego, California

Stefan Christmann / Emperor penguins (Aptenodytes forsteri) silhouetted at sunset, Atka Bay, Queen Maud Land, Antarctica. October.

Jeff Foott / Geyser spouting, Old Faithful, Yellowstone National Park, Wyoming

Andy Rouse / African Lion (Panthera leo) male at sunrise, Masai Mara, Kenya

Staffan Widstrand / Wild Wonders of China / Pere David's deer / Milu (Elaphurus davidianus) male walking in the morning,Hubei Tian'ezhou Milu National Nature Reserve, Shishou, Hubei, China

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) on sea ice at sunrise. Atka Bay, Antarctica. May 2017.

Stefan Christmann / Cameraman Lindsay McCrae on sea ice filming frozen landscape and icebergfor BBC Dynasties Penguin programme. Atka Bay, Antarctica. October 2017.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) standing in drifting snow, silhouetted by sun. Atka Bay, Antarctica. September.

Sven Zacek / Mist over bog with conifers in pre-dawn light. Endla Nature Reserve, Jogevamaa, Central Estonia. June 2016.

Guy Edwardes / Leopard (Panthera pardus) climbing tree, silhouetted against the sun, Kruger National Park, South Africa.

Pedro Narra / Red fox (Vulpes vulpes) silhouetted, Sado Estuary, Portugal. September

Paul Hobson / Wood warbler (Phylloscopus sibilatrix) singing whilst perched on branch, silhouetted at dawn. Sheffield, England, UK. May.

Pal Hermansen / Bear (Ursus arctos) silhouetted at night under starry sky. Finland. August.

Enrique Lopez-Tapia / Man leading Bactrian camels (Camelus bactrianus) in sand dunes, Nubra Valley. Ladakh,,India, September 2018.

Mark Bowler / Aerial shot of the canopy of the Amazon Rainforest at dawn, Rio Tambopata, Madre de Dios, Peru, 2016.

Jack Dykinga / Aurora Borealis reflected in Polar Lake, conifers silhouetted at edge. Near Great Slave Lake, Northwest Territories, Canada. September 2018.

Tui De Roy / Alcedo giant tortoise (Chelonoidis vandenburghi) silhouetted at sunrise, Alcedo Volcano, Isabela Island, Galapagos

Tui De Roy / Marine iguanas (Amblyrhynchus cristatus) silhouettes at sunset, Punta Vicente Roca, Isabela Island, Galapagos

Juergen Freund / Timelapse of a Strangler fig (Ficus virens) at night lit by the moon, known as the Curtain Fig Tree, this individual is over 500 years old, Atherton Tablelands, Wet Tropics World Heritage Area, Queens...

Juergen Freund / Tracking timelapse of stars and the Milky Way seen through the rainforest canopy, Wet Tropics World Heritage Area, North Queensland, Australia.

Juergen Freund / Timelapse of Wallaman Falls at night, lit by the moon, Wet Tropics World Heritage Area, Ingham, Queensland, Australia. 2017.

Juergen Freund / Timelapse of a Scolymia coral (Mussidae) feeding, lit with a UV light, Cairns, North Queensland, Australia.

Remi Masson / Water lilies (Nymphaea alba) lit by the sun's rays, seen under water. Alps, Ain, France. June. Highly commended in the the Underwater World category, GDT European Wildlife Photographer of the Year 201...

Eladio Fernandez / Cienaga de Zapata Swamp at sunset, UNESCO Biosphere Reserve, Cuba, January 2009.

Guy Edwardes / Sunrise through stone at Men-An-Tol, Morvah, Cornwall, England, UK. September 2016.

Guy Edwardes / Sunrise over the Marshwood Vale, West Dorset from Quar Hill, Chideock, Dorset, England, UK. May 2013.

Guy Edwardes / Corfe Castle in morning mist, Corfe, Isle of Purbeck, Dorset, England, UK. September 2013.

Guy Edwardes / Corfe Castle in morning mist, Corfe, Isle of Purbeck, Dorset, England, UK. September 2013.

Guy Edwardes / Wimborne Minster at dawn, Dorset, England, UK. August 2015.

Guy Edwardes / Colmer's Hill in mist, Bridport, Dorset, England, UK. May 2014.

Guy Edwardes / Colmer's Hill at sunrise, Bridport, Dorset, England, UK. May 2014.

Guy Edwardes / Colmer's Hill at sunrise, Bridport, Dorset, England, UK. May 2014.

Guy Edwardes / View from Colmer's Hill, Bridport, Dorset, England, UK. April 2016.

Guy Edwardes / View from Colmer's Hill over countryside with low lying mist, Bridport, Dorset, England, UK. April 2016.

Guy Edwardes / The Drongs sea stacks silhouetted at sunset, Hillswick, Northmavine, Shetland, Shetland Isles, Scotland, UK. August 2014.

Guy Edwardes / The Drongs sea stacks silhouetted at sunset, Hillswick, Northmavine, Shetland, Shetland Isles, Scotland, UK. August 2014.

Guy Edwardes / The Drongs sea stacks silhouetted at sunset, Hillswick, Northmavine, Shetland, Shetland Isles, Scotland, UK. August 2014.

Guy Edwardes / Amethyst deceiver (Laccaria amethystina), New Forest National Park, Hampshire, England, UK. November.

Guy Edwardes / Sunrise over Colmer's Hill, Bridport, Dorset, England, UK. June 2010.

Guy Edwardes / Luskentyre beach with three people wading in water, Isle of Harris, Outer Hebrides, Scotland, UK. June 2012.

Guy Edwardes / Old Man of Storr, Trotternish Peninsula, Isle of Skye, Inner Hebrides, Scotland, UK. January 2014.

Bernard Castelein / Scots pine tree (Pinus sylvestris) at sunrise, Klein Schietveld, Brasschaat, Belgium. September.

Bernard Castelein / Frozen water patterns in Klein Schietveld, Brasschaat, Belgium

Bernard Castelein / Scots pine trees (Pinus sylvestris) in wetlands at dawn with the moon, Klein Schietveld, Brasschaat, Belgium

Bernard Castelein / Scots pine trees (Pinus sylvestris) on island in wetlands, Klein Schietveld, Brasschaat, Belgium

Bernard Castelein / Scots pine tree (Pinus sylvestris) with sunrays at sunrise. Klein Schietveld, Brasschaat, Belgium.

Gavin Hellier / Timelapse of the sun rising over Paris and the Eiffel Tower, France, May 2016. Hellier

Sebastian Kennerknecht / South African Giraffe (Giraffa giraffa giraffa) juvenile at sunset, Greater Makalali Private Game Reserve, South Africa

Avijan Saha/ BIA / Asian Elephant (Elephas maximus) juvenile crossing road at night in headlight of car watched by people on the road, West Bengal, India

Ernie Janes / Pheasant (Phasianus colchicus) male flying to roost at sunset with trees silhouetted in background England, UK. December.

Denis-Huot / Masai giraffe (Giraffa camelopardalis tippelskirchi), female and calf at sunset, with Abdim's storks in flight, Masai-Mara Game Reserve, Kenya

Nick Hawkins / Two Atlantic salmon (Salmo salar) silhouetted against sky, Gaspe Peninsula, Quebec, Canada, October.

Paul Bertner / Rainforest at sunrise, Sani Lodge, Ecuador

Theo Bosboom / Banded demoiselle (Calopteryx splendens) at sunrise, resting in the Common nettle, Roosendaal, the Netherlands. July.

Staffan Widstrand / Praying mantis nymph silhouetted through leaf,. Raja Ampat, Western Papua, Indonesian New Guinea

Theo Bosboom / Giants Causeway at night, lit by full moon. County Antrim, Northern Ireland. Winner of the Landscapes Category of the GDT European Wildlife Photographer of the Year 2017.

Tui De Roy / Flightless cormorant (Phalacrocorax harrisi) silhouetted, Punta Espinosa, Fernandina Island, Galapagos

Guy Edwardes / Puffin (Fratercula arctica) wings spread backlit, Great Saltee Island, County Wexford, Republic of Ireland, June.