Pal Hermansen / Common brimstone (Gonepteryx rhamni) flying in summer meadow amongst Knapweed (Centaurea sp). Hvaler, Ostfold, Norway. July.

SCOTLAND: The Big Picture / Ancient Scots pine tree (pinus sylvestris) with dead low branches, in evening light. Cairngorms National Park, Highlands, Scotland, UK. February 2008.

Neil Aldridge / African elephants (Loxodonta africana) drinking in low evening light at a waterhole, Chobe District, Botswana. Vulnerable species.

Pete Oxford / Whale shark (Rhincodon typus) and scuba divers seen from below, Cenderawasih Bay, West Papua. Indonesia.

Andres M. Dominguez / Orchid (Orchis langei) in sunset light, Sierra de Grazalema Natural Park, southern Spain, May.

Claudio Contreras / Green turtle (Chelonia mydas) with rays of sunlight, Akumal, Caribbean Sea, Mexico, January. Endangered species.

Bence Mate / Wild boar (Sus scrofa) at night with the milky way in the background, Gyulaj, Tolna, Hungary. August. Taken using long exposure with flash at night. Winner of the Mammals Category of the GDT Awards 20...

Juergen Freund / Little red flying fox (Pteropus scapulatus) roosting with baby, Atherton Tablelands, Queensland, Australia. May. Second Place in the Man and Nature portfolio category of the Terre Sauvage Nature Image...

Cyril Ruoso / Verreaux's sifakas (Propithecus verreauxi) in Madagascar ocotillo (Alluaudia procera) trees Berenty, Madagascar.

Konrad Wothe / View up into canopy with sun rays shining through the leaves, lowland rainforest, Panguana Reserve, Huanuco province, Amazon basin, Peru.

Cyril Ruoso / Bengal tiger (Panthera tigris tigris) female resting under tree, Kanha National Park, India. Non-ex

Jack Dykinga / Sunset light on twisted Saguaro cactus (Carnegiea gigantea) Sonoran Desert National Monument, Arizona, USA, April.

Ole Jorgen Liodden / Arctic tern (Sterna Paradisaea) flying, Svalbard, Norway, July.

Markus Varesvuo / Great White Egret (Egretta alba) with water splashing from beak, Hungary January

John Abbott / Low angle view of trees in the Hoh Rainforest, Hall of Mosses Trail, Olympic National Park, Jefferson County, Washington, USA, June.

Andy Trowbridge / Arctic Fox (Alopex / Vulpes lagopus) portrait in early morning light, during moult from grey summer fur to winter white. Dovrefjell National Park, Norway, September.

Daniel Heuclin / Primary forest, New Caledonia.

Daniel Heuclin / Giant Houp (Montrouziera cauliflora), New Caledonia, endemic.

Charlie Summers / Black Bear (Ursus americanus) hunting for spawning Salmon during low tide, Neets Creek estuary, Alaska, July.

Michel Roggo / Brown bear (Ursus arctos beringianus) fishing for Sockeye salmon in the Ozernaya River, Kamtchatka, Russia, August

Ernie Janes / Oystercatcher (Haematopus ostralegus) footprints on sand at low tide, Norfolk, England, UK, November.

Fergus Gill / Whooper Swan (Cygnus cygnus) adult bird in afternoon light. Perthshire, Scotland, March.

Alex Mustard,Alex Mustard / A diver and a hydrothermal chimney. Note that he is obscured by the different temperature waters (from the chimney and the surrounding water) which are unable to mix. The sea water was 4 degrees Celsi...

Michael Pitts / Devil rays (Mobula japonica) viewed from below, South Ari atoll, Maldives.

Franco Banfi,Franco Banfi / Piraputanga (Brycon hilarii) in the main spring Aquario Natural that goes in the Rio Baia Bonita, Bonito, Mato Grosso do Sul, Brazil

Felicity Lanchester,Felicity Lanchester / Cloud iridescence forming above a cumulonimbus cloud, caused by light refraction through individual water droplets and ice crystals at high altitude, Sierra Leone.

Franco Banfi / Manta ray (Manta birostris), San Benedicto, Revillagigedo (Socorro) Islands, Mexico, East Pacific Ocean

Willem Kolvoort / Water lily (Nymphaea rudgeana) leaves viewed from below in a black water creek. Suriname, October.

Yukihiro Fukuda,Yukihiro Fukuda / Japanese fireflies (Luciola cruciata) in flight at night, endemic species, Yaku-shima UNESCO World Heritage Site, Kagoshima, Japan, May

Juergen Freund / Whale shark (Rhincodon typus) near a fishing device called ?Bagan?, a stationary outrigger boat, with a net between outriggers and strong light at night to attract anchovies and scad. The whale sharks...

Juergen Freund / Licuala palms (Licuala ramsayi) with light streaming through the canopy, Daintree, Queensland, Australia

Niall Benvie / Cloud formations lit by low sun. Brechin, Angus, Scotland, December 2012.

Jabruson / African Forest elephant (Loxodonta africana cyclotis) subadult scenting air, blurred motion at dusk, Dzanga Bai, Dzanga-Ndoki National Park, Central African Republic

Floris van Breugel / Tiny thimble jellyfish (Linuche unguiculata) drift with the current in the calm turquiose waters of a lagoon in Ofu, American Samoa. The window of sky is called Snell's Window, caused by refraction of...

Rob Jordan / 2020VISION / Arctic tern (Sterna paradisaea) in flight at twilight using slow shutter speed to accentuate blur and motion in low-key light, Inner Farne, Farne Islands, Northumberland, July.

Ross Hoddinott / Field of ripe wheat with vehicle tracks, in late evening light, near Putford, north Devon, UK, August 2011.

Nick Turner / Man lifting fluorescent light tubes at a recycling centre, Stroud, Gloucestershire, UK, February 2008.

Alex Mustard / 2020VISION / A snorkeller in shallow, sunny waters, Man O War Bay, Jurassic Coast, Dorset, UK, August. Model released.

Erlend Haarberg / Sunlit ridges against volcanic scree beneath a low cloud. Eyjafjallajokull, Iceland, April 2010. Highly commended in Wildlife Photographer of the Year 2011 competition, Wild Places category.

John MacPherson / 2020VISION / Bottlenose dolphin (Tursiops truncatus) two breaching in evening light, Moray Firth, Inverness-shire, Scotland, UK, August, sequence 2/2. Photographer quote: 'A day of overcast cloud cleared in the la...

Diane McAllister / The snow at Horsetail Falls' begins to melt in mid-February, enabling the falls to resume once again. The clear sky and low sun angle ignite the rock wall during sundown. Yosemite National Park, Calif...

Juergen Freund / View of sea surface through a Whip coral forest (Junceella sp) Kimbe Bay, West New Britain, Papua New Guinea

Inaki Relanzon / A snowy mountain peak lit by low crepuscular light before sunset. Ordesa National Park, Pyrenees, Aragon, Spain, February 2009.

Juergen Freund / Light streaming into a shallow cave in a dive site called The Passage, with a diver swimming up towards the foliage visible above. North Raja Ampat, West Papua, Indonesia, February 2010. Model release...

Bruno D'Amicis / Sunset light on Mt. Zdiarska Vidla. Belianske Tatry, Slovakia, November 2007.

Solvin Zankl / Bat expert Dirk Karoske and his team take a captured bat from the net with a dragnet. Stralsund, Germany, June 2009

Floris van Breugel / Ancient Bristlecone Pine tree (Pinus aristata) with sunlight shafts beaming through silhouetted gnarled branches, White Mountains, California, USA. July 2007.

Kevin Schafer / Amazon river dolphin / Boto (Inia geoffrensis) underwater with light above, Rio Negro, Amazonia, Brazil, July

Ross Hoddinott / Porthcothan Bay, evening light and sunset at low tide. Near St Eval, Cornwall, England, UK. April 2010.

Ross Hoddinott / Burgh Island and Burgh Island hotel. Evening light at low tide. Taken from Bigbury Bay (Blue Flag beach) South Devon, UK. September 2009.

Philip Stephen / Trawler in low light on the North Sea, July 2010. Property released.

Ross Hoddinott / Sandymouth bay at low tide, stormy evening light after heavy rainfall, Cornwall, UK. May 2009.

Wild Wonders of Europe / Smit / Wild boar (Sus scrofa) in light morning mist feeding on swamp plants during low water tide in flood plains of the Danube, Gornje Podunavlje Special Nature Reserve, Serbia, June 2009

Wild Wonders of Europe / Damschen / Lenzer Wische with light low lying mist at dawn, Elbe Biosphere Reserve, Lower Saxony, Germany, July 2008

Michael Durham / Douglas Fir Glow-worm (Pterotus obscuripennis) displaying bioluminescence at night, Tillamook State Forest, Oregon

Colin Monteath / Mt Makalu (8,462 meters) and Mt Chomolonzo (7,540 meters) bathed in dawn light, reflected in small lake, Khama Valley, Tibet

Norbert Wu / California Sea Lion (Zalophus californianus) playing in shallow water, Channel Islands, California

Flip Nicklin / Giant Kelp (Macrocystis pyrifera) forest, Channel Islands National Park, California

Martin Dohrn / The Southern Cross star constellation shining above Ballenites trees in Masai Mara, Kenya. Starlight image intensifier camera image taken with no artificial light.

Martin Dohrn / Lightning at night over Masai Mara Kenya, East Africa. Starlight image intensifier camera image taken with no artificial light

Martin Dohrn / Bioluminescence - pool with Snook, Snapper and a Stingray fish, Bio Bay, Vieques Island, Puerto Rico. Central America. Starlight image intensifier camera image taken with no artificial light.

Martin Dohrn / Holding head low, a Gazelle watches lion pride walk by. Serengeti NP, Ngorongoro Conservation Area, Tanzania. Image taken at night using 'Starlight Camera' technology and infra red light.

Dave Watts / Platypus swimming underwater {Ornithorhynchus anatinus} Australia. Digital composite.

Bernard Castelein / American oak (Quercus rubra) trees in avenue, sun shining through. Peerdsbos, Brasschaat, Belgium. November 2013.

Shane Gross / Caribbean reef shark (Carcharhinus perezi) and Horse eye jack (Caranx latus) shoal silhouetted underneath catamaran boat, Bahamas.

Shane Gross / Caribbean reef shark (Carcharhinus perezi) swimming over coral reef, silhouetted. Caribbean Sea off Gardens of the Queen National Park, Cuba.

Heather Angel / Alexander's rhubarb (Rheum alexandrae) and Dwarf purple rhododendron (Rhododendron impeditum) in wet valley with low cloud. Large bracts on rhubarb protect tiny flowers from UV light. Near Lake Ka...

Jeff Foott / California Poppy (Eschscholzia californica) flowers, super bloom, Antelope Valley, California

Franco Banfi / Scuba diver between trees seen from below the surface surrounding Lagoa Misteriosa, a sink hole in limestone region, Serra da Bodoquena (Bodoquena Mountain Range), Bonito, Mato Grosso do Sul, Brazil

Franco Banfi / Scuba diver between trees seen from below the surface surrounding Lagoa Misteriosa a sink hole in limestone region, Serra da Bodoquena (Bodoquena Mountain Range), Bonito, Mato Grosso do Sul, Brazil

Jack Dykinga / Rays of low sunlight through Douglas fir (Pseudotsuga menziesii) trees in early morning fog, Redwood State and National Parks, California, USA. May.

Jack Dykinga / Rays of low sunlight through Douglas fir (Pseudotsuga menziesii) trees in early morning fog, Redwood State and National Parks, California, USA. May.

Jack Dykinga / Rays of low sunlight through Douglas fir (Pseudotsuga menziesii) trees in early morning fog, Redwood State and National Parks, California, USA. May.

Ashley Cooper / Old man with a low energy light bulb which he can use to power house with solar charged batteries from WWF project to supply electricity to a remote island in the Sunderbans, Ganges Delta, India. Dece...

Ashley Cooper / Boy holding a solar lantern, charged from the solar panelsfrom WWF project to supply renewable electricity to a remote island in the Sunderbans, Ganges, India. December 2013

Ashley Cooper / Young boy doing homework in house lit by electricity from battery. Batteries are solar charged from WWF project to supply electricity to a remote island in the Sunderbans, Ganges Delta, India. Decembe...

Ashley Cooper / Man with new solar lanterns from WWF project to supply electricity to a remote island in the Sunderbans, Ganges Delta, India. December 2013

Ashley Cooper / Man with new solar lanterns from WWF project to supply electricity to a remote island in the Sunderbans, Ganges Delta, India. December 2013

Ashley Cooper / Drought impacted lake in Yosemite National Park, California, USA. October 2014.

Ashley Cooper / Electric vehicle at a recharging station on the street in Berkeley Square, London, UK. May 2012.

Ashley Cooper / Electric vehicle at a recharging station on the street in Berkeley Square, London, England, UK. May 2012.

Richard Robinson / Grey Unicornfish (Naso caesius) and Sleek Unicorn fish (Naso hexacanthus ) school at Banc Lorne, New Caledonia.

Jen Guyton / Speckled tree snake (Imantodes inornatus) on a Heliconia flower, La Selva Biological Station, Costa Rica.

Melvin Grey / Lapwing (Vanellus vanellus) adult female with chick, rough grazing meadow in low evening light. North Uist, Scotland, UK, May.

Sandra Bartocha / Rocks with splashes from the surf with low sun, Stockholm Archipelago, Sweden

Sandra Bartocha / Panoramic of Oak tree (Quercus robur) with low sun, Oland, Sweden, May.

Juergen Freund / Purple crown jellyfish (Netrostoma setouchina) in the shallow waters, Nukubati Island Resort, Macuata Province, Fiji, South Pacific

Frederique Olivier / Close-up of a Light-mantled sooty albatross (Phoebetria palpebrata) preening, Macquarie Island, Sub-Antarctic Australia.

Frederique Olivier / Close-up of a Light-mantled sooty albatross (Phoebetria palpebrata) with droplets of water on its back, Macquarie Island, Sub-Antarctic Australia.

Frederique Olivier / Close-up of a Light-mantled sooty albatross (Phoebetria palpebrata) with droplets of water on its head, Macquarie Island, Sub-Antarctic Australia.

Frederique Olivier / Close-up of a Light-mantled sooty albatross (Phoebetria palpebrata) with droplets of water on its head, Macquarie Island, Sub-Antarctic Australia.

Frederique Olivier / Close-up of a Light-mantled sooty albatross (Phoebetria palpebrata) building its nest, Macquarie Island, Sub-Antarctic Australia.

Frederique Olivier / Close-up of a Light-mantled sooty albatross (Phoebetria palpebrata) building its nest, Macquarie Island, Sub-Antarctic Australia.

Ingo Arndt / Aurora borealis / Northern lights over boreal forest near Denali National Park, Alaska, USA, March 2013.