Alex Mustard / Split level image of Leather corals (Sarcophyton sp.) flourishing in shallow water close to uninhabited coral islands. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean...

Doug Perrine / Pygmy shark (Euprotomicrus bispinatus) portrait, captive. Kona, Hawaii. Central Pacific Ocean. Digitally manipulated image.

Suzi Eszterhas / Sea Otter (Enhydra lutris) raft, Monterey Bay, California, digitally manipulated

Ernie Janes / Abstract photo of Tower Bridge, with blurred motion red bus, London, June 2013. Digitally manipulated.

Alex Mustard / Blue shark (Prionace glauca) near the surface of the English Channel. Penzance, Cornwall, England, British Isles. North East Atlantic Ocean, August. Digitally manipulated to improve composition see 1...

Ernie Janes / Robin (Erithacus rubecula) in snow, Titchwell, Norfolk, January. Image digitally manipulated to add more snow.

Pete Oxford / Amazon Tree Boa (Corallus hortulanus) in rainforest, Iwokrama Reserve, Guyana, digitally manipulated image

Alex Mustard / Split level image of Caribbean reef shark (Carcharhinus perezi) breaking surface at dusk. Grand Bahama, Bahamas. Digitally manipulated image.

Pete Oxford / Common Tree Boa (Corallus hortulanus) coiled around branch, Iwokrama Rainforest Reserve, Guyana, manipulated image

Pete Oxford / Common Tree Boa (Corallus hortulanus) climbing in tree, Iwokrama Rainforest Reserve, Guyana, manipulated image

Pete Oxford / Common Tree Boa (Corallus hortulanus) wrapped around branch, Iwokrama Rainforest Reserve, Guyana, manipulated image

Doug Perrine / Dolphin fish / Mahi mahi / Dorado (Coryphaena hippurus) close up, off Isla Mujeres, near Cancun, Yucatan Peninsula, Mexico, Caribbean Sea (digitially manipulated)

Murray Cooper / Sparkling Violet-ear (Colibri coruscans) hummingbird males fighting, Ecuador, digitally manipulated

Niall Benvie / Photograph of flowering Burnet saxifrage (Pimpinella saxifraga) manipulated and text added to represent illustrated book page. Scotland, UK, July

Niall Benvie / Photograph of flowering Restharrow (Ononis sp)manipulated and text added to represent illustrated book page. Scotland, UK, July

Niall Benvie / Photograph of Hairy St John's wort (Hypericum hirsutum) manipulated and text added to represent illustrated book page. Scotland, UK, July

Niall Benvie / Photograph of Blackthorn blossom (Prunus spinosa) manipulated and text added to represent illustrated book page. Scotland, UK, July

Stephen Dalton / Barn owl (Tyto alba) in flight, flying down to catch rat prey, UK, manipulated image (controlled conditions rat protected by sheet of perspex)

Doug Perrine / Atlantic sailfish {Istiophorus albicans} hits sardine with bill while feeding on bait ball of Spanish sardines / gilt sardine / pilchard / round sardinella {Sardinella aurita} off Yucatan Peninsula, M...

Doug Perrine / Atlantic sailfish {Istiophorus albicans} attacking bait ball of Spanish sardines / gilt sardine / pilchard / round sardinella {Sardinella aurita} off Yucatan Peninsula, Mexico, Caribbean Sea, Digitall...

Doug Perrine / Atlantic sailfish {Istiophorus albicans} attacking bait ball of Spanish sardines / gilt sardine / pilchard / round sardinella {Sardinella aurita} off Yucatan Peninsula, Mexico, Caribbean Sea, Digitall...

Doug Perrine / False killer whale ( Pseudorca crassidens ) with mouth open, showing large conical teeth, captive, aquarium, from Indo-pacific, digitally manipulated

Doug Perrine / Young Sperm whale calf (Physeter macrocephalus) Endangered, Kona, Hawaii, Central Pacific Ocean digitally manipulated

Doug Perrine / Northern / lined / Atlantic seahorse (Hippocampus erectus) male giving birth - expelling fry from pouch - young fry then float to the water's surface; captive, digitally manipulated

Doug Perrine / Bottlenose dolphins, captive {Tursiops truncatus} Honduras. digitally manipulated (Non-ex).

Doug Perrine / Sociable wild Bottlenose dolphin {Tursiops truncatus} Belize - Honey (Non-ex).

Doug Perrine / Subadult Pelagic thresher shark {Alopias pelagicus} Philippines, Pacific Ocean (Non-ex).

Alex Mustard / Blue whale (Balaenoptera musculus) diving in Indian Ocean, Sri Lanka. Digitally manipulated, left side extended.

Alex Mustard / Blue tang (Acanthurus coeruleus) with Bluehead (Thalassoma bifasciatum) juveniles at cleaning station on coral reef. George Town, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea. Digi...

Alex Mustard / Mirrored portrait of a Weedy scorpionfish (Rhinopias frondosa). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea. Digitally manipulated (the face is reflected down mid-line).

Kim Taylor / Common earwig (Forficula auricularia) closeup of head. Surrey, England, September. Digitally manipulated.

Kim Taylor / Woodlouse (Oniscus asellus) close up of head. Surrey, England, September. Digitally manipulated.

Wim van den Heever / Zebra (Equus quagga) leaping during stampede, Serengeti, Tanzania. Vignette added and right edge expanded.

Markus Varesvuo / Spotless starling (Sturnus unicolor) Spain, December. Digitally manipulated, one minor part of second bird removed

Markus Varesvuo / Redwing (Turdus iliacus) perched amongst berries, Helsinki, Finland, November. (Digitally manipulated - one branch removed)

Markus Varesvuo / Pine grosbeak (Pinicola enucleator) female feeding on berries, Helsinki, Finland, November. Digitally manipulated - One branch removed

Markus Varesvuo / White-backed woodpecker (Dendrocopus leucotos) perched on branch, Helsinki, Finland, January, digitally manipulated - one branch removed

Markus Varesvuo / King Eider (Somateria spectabilis) males and a female in flight over water, Norway, April. Digitally manipulated, One bird removed

Markus Varesvuo / Griffon vulture (Gyps fulvus) perched on rock, vocalising, Spain, December, digitally manipulated

Markus Varesvuo / Eurasian Jackdaw (Corvus monedula) perched on stones, Finland, December, digitally manipulated

Markus Varesvuo / Coal tit (Periparus ater) feeding on peanuts from bird feeder, Finland, digitally manipulated

Markus Varesvuo / Flock of Common myna (Acridotheres tristis) in flight, Oman, March, digitally manipulated

Alex Mustard / Portrait of female Scalefin anthias (Pseudanthias squamipinnis) feeding on zooplankton in open water. Ras Katy, Sinai, Egypt. Gulf of Aqaba, Red Sea. Manipulated - canvas rotation and extension. No ch...

Alex Mustard / Giant cuttlefish (Sepia apama) blows water out through its syphon to jet backwards through sea grass meadow. Manly, Sydney, New South Wales, Australia, digitially manipulated image

Dave Watts / Greater flamingos (Phoenicopterus ruber) in flight at dawn, Camargue, France, digitally manipulated

Dave Watts / Southern cassowary (Casuarius casuarius) adult male with three chicks, World Heritage National Park rainforest of the Wet Tropics, north Queensland, Australia, digitally manipulated

Dave Watts / Southern cassowary (Casuarius casuarius) adult male crossing road near road warning sign, World Heritage National Park rainforest of the Wet Tropics, north Queensland, Australia, digital composite

Alex Mustard / Smooth starlet coral (Siderastrea sidera) close focus wide angle view of a massive colony. This photo is not digital manipulated, but taken with a fisheye lens placed very close to the coral with flas...

Alex Mustard / Close focus wide angle view of a colony of Great star coral (Montastrea cavernosa). This photo is not digital manipulated, but taken with a fisheye lens placed very close to the coral with flashes aim...

Alex Mustard / Brain coral (Pseudodiploria strigosa) close focus wide angle view. Photograph is not digital manipulated, but taken with a fisheye lens placed very close to the coral with flashes aimed to illuminate...

Pete Oxford / Timber rattlesnake (Crotalus horridus) yellow morph near hibernation den, Northern Georgia, USA, manipulated

Pete Oxford / Timber rattlesnakes (Crotalus horridus) black and yellow morphs, near hibernation den, Northern Georgia, USA, manipulated

Pete Oxford / Timber rattlesnake (Crotalus horridus) black morph near hibernation den, Northern Georgia, USA, manipulated

Pete Oxford / Timber rattlesnakes (Crotalus horridus) black and yellow morphs near hibernation den, Northern Georgia, USA, manipulated

Pete Oxford / Eastern Narrowmouth Toad (Gastrophryne carolinensis) The Orianne Indigo Snake Preserve, Telfair County, Georgia, USA, manipulated image

Pete Oxford / Southern Toad (Bufo / Anaxyrus terrestris) The Orianne Indigo Snake Preserve, Georgia, USA, digitally manipulated image

Hanne & Jens Eriksen / Three Brown pelicans (Pelecanus occidentalis) in flight, digitally manipulated, Texas, USA, January

Ernie Janes / Flock of Dunlin (Calidris alpina) flying past a rising moon, Norfolk, UK (Digitally manipulated)

Pete Oxford / Timber Rattlesnake (Crotalus horridus) black morph, native to the eastern United States, Digitally Manipulated

Pete Oxford / Hippopotamus (Hippopotamus amphibius) underwater showing teeth, Lowveld, South Africa, digitally manipulated

Markus Varesvuo / Bearded vulture (Gypaetus barbatus) subadult landing in snow, Spain, November, digitally manipulated

Pete Oxford / Amazon Tree Boa (Corallus hortulanus) coiled on rainforest branch, Iwokrama Reserve, Guyana, digitally manipulated image

Pete Oxford / Amazon Tree Boa (Corallus hortulanus) in rainforest, Iwokrama Reserve, Guyana, digitally manipulated image

Doug Perrine / Bryde's whale, (Balaenoptera brydei / edeni) Striped marlin (Tetrapturus audax) and California sea lions (Zalophus californianus) off Baja California, Mexico,Eastern Pacific Ocean, November. (digitall...

Suzi Eszterhas / Bengal Tiger (Panthera tigris tigris) two year old cubs playing, Bandhavgarh National Park, India, digitally manipulated

Suzi Eszterhas / Hog Deer (Axis porcinus) female, Kaziranga National Park, India, digitally manipulated

Philippe Clement / Reflection in pond of Nessie, the Loch Ness monster, outside the Loch Ness Exhibition Centre at Drumnadrochit, Scotland, UK May 2010 - digitally manipulated

Pete Oxford / Common Tree Boa (Corallus hortulanus) in branch, Iwokrama Rainforest Reserve, Guyana, manipulated image

Pete Oxford / Common Tree Boa (Corallus hortulanus) wrapped around branch, Iwokrama Rainforest Reserve, Guyana, manipulated image

Hanne & Jens Eriksen / Broad-billed sandpiper {Limicola falcinellus} three in flight over water, Oman, January, digitally manipulated

Doug Perrine / Dolphin fish / Mahi mahi / Dorado (Coryphaena hippurus) off Isla Mujeres, near Cancun, Yucatan Peninsula, Mexico, Caribbean Sea (digitally manipulated)

Niall Benvie / Photograph of Alpine penny-cress (Thlaspi caerulescens) manipulated and text added to represent illustrated book page. Scotland, UK, July

Niall Benvie / Photograph of Alpine rock-cress (Arabis alpina)manipulated and text added to represent illustrated book page. Scotland, UK, July

Niall Benvie / Male and female Stag beetle (Lucanus cervus) Photograph with text inserted, and manipulated to appear as book illustration, England, UK

Doug Perrine / Live deep-sea anglerfish / black seadevil (Diceratias pileatus) captive, showing bioluminescent lure and razor sharp teeth, specimen brought up from a depth of 3,300 feet (1000m) in a water intake pi...

Doug Perrine / Live deep-sea anglerfish / black seadevil (Diceratias pileatus) captive, showing bioluminescent lure and razor sharp teeth, specimen brought up from a depth of 3,300 feet (1000m) in a water intake pi...

Doug Perrine / Live deep-sea anglerfish / black seadevil (Diceratias pileatus) captive, showing bioluminescent lure and razor sharp teeth, specimen brought up from a depth of 3,300 feet (1000m) in a water intake pi...

Doug Perrine / Close up of mouth of Deep-sea anglerfish / black seadevil (Diceratias pileatus) captive, showing needle-sharp teeth; specimen brought up from a depth of 3,300 feet (1000m) in a water intake pipe at N...

Murray Cooper / Rufous-crested Coquette (Lophornis delattrei) hummingbird male, Colombia, digitally manipulated

Hanne & Jens Eriksen / Black tailed godwit (Limosa limosa) in flight, Iceland, June (Digitally manipulated)

Suzi Eszterhas / Topi (Damaliscus lunatus) trio in grassland, Masai Mara National Reserve, Kenya, Digitally Manipulated

Hanne & Jens Eriksen / Caspian tern {Hydroprogne caspia} two in flight, digitally manipulated, USA

Hanne & Jens Eriksen / Black skimmer {Rhynchops niger}, three in flight over water, digitally manipulated, Texas, USA

Wild Wonders of Europe / Varesvuo / European Roller (Coracias garrulus) pair in courtship, Pusztaszer, Hungary, May 2008, Digitally manipulated - One part of a branch removed

Doug Perrine / Atlantic sailfish {Istiophorus albicans} attacking bait ball of Spanish sardines / gilt sardine / pilchard / round sardinella {Sardinella aurita} off Yucatan Peninsula, Mexico, Caribbean Sea. Digitall...

Doug Perrine / Atlantic sailfish {Istiophorus albicans} off Yucatan Peninsula, Mexico, Caribbean Sea. Digitally manipulated

Doug Perrine / Striped marlin {Tetrapturus audax} feeding on baitball of Sardines {Sardinops sagax} off Baja California, Mexico, Digitally manipulated

Doug Perrine / Striped marlin {Tetrapturus audax} feeding on baitball of Sardines {Sardinops sagax} off Baja California, Mexico, Digitally manipulated

Doug Perrine / With a deadly swipe of its bill, a Striped marlin {Tetrapturus audax} dispatches a Sardine / Pilchard {Sardinops sagax} and prepares to swallow it, after chasing it out of a bait ball, off Baja Califo...

Doug Perrine / Porbeagle shark (Lamna nasus) captive, Nova Scotia, Canada, image digitally manipulated

Doug Perrine / Porbeagle shark (Lamna nasus) captive, Nova Scotia, Canada, image digitally manipulated - note white patch on rear edge of dorsal fin and double keel on caudal (tail) fin that are diagnostic of this...

Doug Perrine / Porbeagle shark (Lamna nasus) captive, Nova Scotia, Canada, image digitally manipulated

Doug Perrine / Porbeagle shark (Lamna nasus) captive, Nova Scotia, Canada, image digitally manipulated

Doug Perrine / Porbeagle shark (Lamna nasus) captive, Nova Scotia, Canada, image digitally manipulated

Doug Perrine / Green sea turtle (Chelonia mydas) hatchling swimming, Endangered, Caribbean, captive, digitally manipulated

Doug Perrine / Tiger shark (Galeocerdo cuvier) lunging for bait, North Shore, Oahu, Hawaii, USA, Central Pacific Ocean (digitally manipulated)