Martin Dohrn / Group of lions (Panthera leo) at night, Masai Mara, Kenya. Photographed using 'Starlight Camera' technology without artificial lighting.

Claudio Contreras / American Flamingo (Phoenicopterus ruber) juveniles following adults, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, August.

Alex Mustard / A school of lesser devil rays (Mobula hypostoma) flying through sunbeams as they feed. Isla Mujeres, Quintana Roo, Yucatan Peninsula, Mexico. Caribbean Sea.

Paul D Stewart / Lithograph with contemporary colouring, continental version of the wallchart produced by Benjamin Waterhouse Hawkins for the Department of Science and Art 1862. It is entitled 'Enaliosauria, or marine...

Jabruson / Mozambican children with captive Yellow Baboon youngster (Papio cynocephalus). Young animal caught during troop crop raiding. Pemba to Montepuez highway, north-eastern Mozambique, November 2011. Wildl...

Juan Carlos Munoz / Aerial view of flock of Flamingos flying over the salt lake, Lake Natron, Rift Valley, Tanzania, August 2009

Juan Carlos Munoz / Common carp (Cyprinus carpio) in lake, Plitvice Lakes National Park, Lika, Croatia, Europe, October 2011

Erlend Haarberg / Eight Rock ptarmigans (Lagopus muta) with snow blowing in wind, Norway, February

Mike Read / Cranes (Grus grus), imprinted from hatching to fly with a microlight. Hampshire County Show near Brockenhurst, Hampshire, England, July 2008.

Paul Hobson / Birch Sawfly (Cimbex femoratus) feeding on a leaf. August, UK.

Laurie Campbell / Mute swans (Cygnus olor) newly hatched cygnets on pond, Yetholm Loch Scottish Wildlife Trust Reserve, Roxburghshire, Scotland, May

Todd Mintz / Sockeye salmon (Oncorhynchus nerka) split level view of annual spawning run, Adams River, British Columbia, Canada, October

Luiz Claudio Marigo / Family of Southern coatis (Nasua nasua) in Atlantic Rainforest, Tijuca National Park, Rio de Janeiro City, Southeastern Brazil.

Alex Hyde / African moon moth (Argema mimosae) digital composite sequence, showing wings expanding after emerging from cocoon, photographed against a white background, orginating from Africa. Captive.

Pedro Narra / Common dolphins ( Delphinus delphis) swimming below the surface, Atlantic Ocean, Portugal, October

Pedro Narra / A pod of Bottlenose dolphins (Tursiops truncatus) swimming into the Sado river, Portugal, July

Chadden Hunter / King penguins (Aptenodytes patagonicus) walk along beach, South Georgia. Taken on location for BBC Frozen Planet series, 2008

Chadden Hunter / Emperor penguins (Aptenodytes forsteri) males dive through openings in the sea ice to take their first swim in over 4 months as they have spent the bleak winter incubating their eggs. Cape Washington,...

Simon Colmer / Honey bee (Apis mellifera) returning to hive and communicating nectar source to other workers, Sussex, UK

Simon Colmer / Honey bee (Apis mellifera) workers exchanging food - known as trophallaxis, Sussex, UK

John Abbott / Tourists watching large colony of Marine Iguanas (Amblyrhynchus cristatus) Punta Espinosa on Fernandina Island, Galapagos, August 2010

Juan Carlos Munoz / Jersey tiger moth (Euplagia quadripunctaria) large numbers flying to and resting on Sweetgum trees (Liquidambar orientalis) in the Petaloudes Valley, Rhodes island, Greece, August 2011

Kristel Richard / An Indian cowboy has lost his balance from a bronc or wild paint horse during the All Indian Rodeo, at the annual Indian Crow Fair, at Crow Agency, near Billings, Montana, USA, August 2011, Sequence 3...

Theo Allofs / African Elephant (Loxodonta africana) herd crossing desert, Skeleton Coast, Namib Desert, Namibia

Ben Osborne / African elephants (Loxodonta africana) bulls crowd into an emptying waterhole to drink, Savuti, Botswana. Taken on location for BBC Planet Earth series, 2005

Konrad Wothe / Southern Cassowary (Casuarius casuarius) female walking near tourists on beach, Moresby Range NP, Queensland, Australia, November 2010

Sven Zacek / Roe Deer (Capreolus capreolus) herd travelling through snow and fog. Virumaa, Estonia, Europe, March.

Sven Zacek / Geese on their spring migration above dawn lit clouds. Estonia, Europe, May.

Sven Zacek / Small herd of Roe Deer (Capreolus capreolus) looking back at the photographer against forested background. Virumaa, Estonia, Europe, February.

Sven Zacek / Herd of Roe Deer (Capreolus capreolus) on the horizon with another out-of-focus in foreground. Virumaa, Estonia, Europe, February.

Sven Zacek / Fish seen through water and the reflections of trees. Jogeva maakond, Estonia, Europe, February.

Andy Rouse / Walrus (Odobenbus rosmarus) colony discharging gas on beach. Svalbard, Norway, August 2011.

Andy Rouse / Groups of Burchell's Zebra (Equus quagga) in grassland seen from balloon. Masai Mara, Kenya, Africa. (non-ex).

Ernie Janes / Five newly hatched chicks in a row

Ernie Janes / Gulls on the beach at sunset, The Wash, Hunstanton, Norfolk, UK, September 2011

Ernie Janes / House sparrows (Passer domesticus) perched on building, Morocco, March

Richard Du Toit / Low level view of domestic Ostrich (Struthio camelus), Overberg, Southern Cape, South Africa August

Florian Möllers / Chinese fisherman on his raft with Great Cormorant (Phalacrocorax carbo sinensis), silhouetted against the karst hills at the Li River. Yangshuo, Guangxi, China, November.

Fabrice Cahez / Group of Wild boar (Sus scrofa) silhouetted, Vosges, France, June

Andrew Parkinson / Mallard (Anas platyrhynchos) line of ducklings follow their mother through early morning mist, Derbyshire, UK, June.

Andrew Parkinson / Lapwing (Vanellus vanellus) group roosting in the lagoons at Titchwell RSPB Reserve, Norfolk, UK, October.

Sandesh Kadur / Great-tailed Grackle (Quiscalus mexicanus) roosting on electric lines along Highway 77, Brownsville, Texas, USA. March

Sandesh Kadur / Greater adjutant storks (Leptoptilos dubius) at a landfill dite near the city of Guwahati, Assam, India, Critically Endangered

B&S Draker / Familiar Bluet Damselflies (Enallagma civile) males perched along a stem. Sinton, Corpus Christi, Coastal Bend, Texas, USA, April.

Grzegorz Lesniewski / Herd of Carpathian Pony / Hucul (Equus caballus) in snow. Bieszczady National Park, the Carpathians, Poland, March.

Piotr Naskrecki / Cross Frog (Oreophryne sp) male protecting his clutch of eggs, Papua New Guinea

Bence Mate / Dalmatian pelicans (Pelecanus crispus) low angle perspective of open bills, Lake Kerkini, Greece, February. Highly Commended, 2011 Wildlife Photographer of the Year competition.

Doug Perrine / Green sea turtle (Chelonia mydas) being cleaned of algae by Yellow tangs / surgeonfish (Zebrasoma flavescens), Puako, Kona, Hawaii, USA, June, Endangered species

Bence Mate / Leaf cutter ants (Atta sp) female worker ants carry pieces of fern leaves to nest, reflected in water, Costa Rica, December. Winner, Eric Hosking award, 2011 Wildlife Photographer of the Year competit...

Bence Mate / Dalmatian pelican (Pelecanus crispus) low angle perspective of open bill, Lake Kerkini, Greece, February. Winner, Eric Hosking award, 2011 Wildlife Photographer of the Year competition

Markus Varesvuo / Flock of Crane (Grus grus) flying overhead with a cliff in the foreground. Spain, March.

Philippe Clement / Grey Heron (Ardea cinerea) footprints in snow. Belgium, December 2010.

Ann & Steve Toon / Domestic sheep (Ovis aries) flock of Northumberland blackface sheep in snow, Tarset, Hexham, Northumberland, UK, November 2010

Jeff Rotman / Mattanza fishermen landing Giant Bluefun tuna (Thunnus thynnus) using a technique in which a maze of nets are set, leading to a central pool, Favignana, Italy, May / June 1990

Doug Perrine / Reef manta rays (Manta alfredi formerly Manta birostris) filter feeding on plankton among Silverside fish that are also feeding on the plankton, Hanifaru Bay, Hanifaru Lagoon, Baa Atoll, Maldives, Ind...

Jeff Vanuga / Wild Horses (Equus caballus) grazing in evening light. Pryor Mountain, border of Montana and Wyoming, USA, June.

Eric Baccega / Black-legged Kittiwake (Rissa tridactyla) on ice. Floe edge, Arctic Bay, Nunavut, Canada, April.

Troels Jacobsen/Arcticphoto / A group of Rockhopper Penguins (Eudyptes chrysocome) under long coastal grasses. The world population of Northern Rockhopper has declined by 90% in the past 60 years. Nightingale Island, Tristan da Cu...

Juergen Freund / School of Bigeye jacks / Trevally (Caranx sexfasciatus). Misool, Raja Ampat, West Papua, Indonesia

Juergen Freund / Schooling Banggai cardinalfish in anemone tentacles. (Pterapogon kaudemi). Lembeh Strait, North Sulawesi, Indonesia

Juergen Freund / Nudibranchs (Hypselodoris / Chromodoris bullocki) mating and laying eggs, Lembeh Strait, North Sulawesi, Indonesia

Juergen Freund / Dock worker carrying Yellowfin tuna ashore for weighing, Jacana tuna fish landing, Puerto Princesa, Palawan, Philippines, April 2009

Juergen Freund / Oceanic sunfish (Mola mola) with bannerfish and angelfish cleaning it. Taken at 42 meters. Nusa Penida, Lesser Sunda Islands, Indonesia,

Tony Phelps / Scorpion (Uroplectes lineatus / carinatus) female carrying hatchlings on its back; a rare example of parental care in invertebrates. De Hoop Nature Reserve, Western Cape, South Africa, January.

Colin Varndell / Red Deer (Cervus elaphus) hinds silhouetted in grassland. Richmond Park, London, UK, October.

Anup Shah / African elephant (Loxodonta africana) group leaving a waterhole - wide angle perspective. Masai Mara National Reserve, Kenya. March

Anup Shah / African elephant (Loxodonta africana) group drinking from waterhole - wide angle perspective. Masai Mara National Reserve, Kenya. March

Anup Shah / Common / Plains Zebra (Equus quagga burchellii) herd running away from the Mara River - wide angle perspective, Masai Mara National Reserve, Kenya. September 2009.

Anup Shah / Flock of White-backed vultures (Gyps africanus) feeding on a carcass, wide angle perspective. Masai Mara National Reserve, Kenya. September

Anup Shah / Thomson's gazelles (Eudorcas thomsonii) following a herd of Wildebeest (Connochaetes taurinus) as they cross the Mara River, wide angle perspective, Masai Mara National Reserve, Kenya. September

Anup Shah / Eastern White-bearded Wildebeest (Connochaetes taurinus) herd running - wide angle perspective, Masai Mara National Reserve, Kenya. August 2009.

Anup Shah / Eastern White-bearded Wildebeest (Connochaetes taurinus) herd crossing the Mara River, close up of legs - wide angle perspective. Masai Mara National Reserve, Kenya. September 2009.

Edwin Giesbers / Five Mallard (Anas platyrhynchos) walking in a line across snow. Three males and two females. The Netherlands, January.

Martin Camm / Carwardine / Illustration of Common bottle-nosed / Bottlenose dolphin (Tursiops truncatus) and Indo-Pacific dolphin (T. aduncus) breach and dive sequence in profile (Wildlife Art Company).

Chris Shields / Illustration of wildlife at African waterhole: Water buffalo (Syncerus caffer),gazelle,African lion (Panthera leo),African elephant (Loxodonta sp), weaver birds (Ploceidae).

Martin Camm / Carwardine / Illustration of magnetic fields and dolphins stranding (Wildlife Art Company).

Sergey Gorshkov / Migrating Salmon (Salmonidae) Lake Kuril, Kamchatka, Far East Russia, August

Tim Laman / Marine Iguanas (Amblyrhynchus cristatus) on the south coast of Espanola Island, Galapagos, Vulnerable species

Georgette Douwma / Lyrtetail anthias (Pseudanthias squamipinnis) in water column with Peach anthias (Pseudanthias dispar) in background. Indonesia, October

David Noton / Woman and man with livestock walking along path, Isla del Sol, Lake Titicaca, Bolivia, December 2009

Sue Flood / Bluefin tuna {Thunnus thynnus} captive in fish farm, Japan, 2000

Yva Momatiuk & John Eastcott / Sockeye Salmon (Oncorhynchus nerka) group swimming upstream between forested banks during spawning run, Adams River, Roderick Haig-Brown Provincial Park, British Columbia, Canada

Yva Momatiuk & John Eastcott / Sockeye Salmon (Oncorhynchus nerka) group swimming upstream between forested banks during spawning run, Adams River, Roderick Haig-Brown Provincial Park, British Columbia, Canada

Kim Taylor / Wood Ant (Formica rufa) workers on top of their nest synchronise ejection of formic acid droplets to maximise deterrent effect on potential predator, UK. Commended in the Audubon Society of Greater De...

Bernard Castelein / Cinereous / European Black Vulture (Aegypius monachus) in dispute with an Eurasian Griffon Vulture (Gyps fulvus), alongside a Himalayan Griffon (Gyps himalayensis) and Long-billed Vulture (Gyps indicu...

Anup Shah / African elephant (Loxodonta africana) investigating with its trunk, Masai Mara National Reserve, Kenya. March

Anup Shah / African elephant herd (Loxodonta africana) spraying mud with trunk, Masai Mara National Reserve, Kenya. December

Anup Shah / Common or Plains zebra (Equus quagga burchellii)crossing the Mara River. Masai Mara National Reserve, Kenya, September 2009

Ernie Janes / Household Cavalry exercising their horses on Holkham Beach, Norfolk, UK, July 2008

Ernie Janes / Free range Domestic pig (Sus scrofa domesticus) four piglets at four weeks, UK, July 2010

David Tipling / Pied wagtails (Motacilla alba yarrellii) silhouetted against the moon, roosting in tree in shopping centre. Kent, UK, December

Laurent Geslin / Herring gull (Larus argentatus) swooping down to feed on chips in a busy seaside resort, France

ASHISH & SHANTHI CHANDOLA / Rabari caravan with women leading camels loaded up for the journey, Kutch, Gujarat, India, August 2009

Andrew Parkinson / Pair of Greylag geese (Anser anser) and chicks spotlit by dawn sunlight near a small weir. Nottinghamshire, UK, May (non-ex)

Michel Poinsignon / Flock of Mute swans (Cygnus olor) in winter, snowing, Lorraine, France, February

Alex Mustard / Group of Manta rays (Manta birostris) feeding together on plankton in a shallow lagoon. Hanifaru Lagoon, Baa Atoll, Maldives. Indian Ocean.