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Close-up of a Meadow brown butterfly (Maniola jurtina) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, England, UK, June.
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Nick Upton / Close-up of a Meadow brown butterfly (Maniola jurtina) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, England, UK, June.
Juvenile Goldenrod spider (Misumena vatia) camouflaged on a Common hogweed (Heracleum sphondylium) flowerhead, reacting to and catching a fly (Muscidae), Wiltshire, England, UK, June.
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Nick Upton / Juvenile Goldenrod spider (Misumena vatia) camouflaged on a Common hogweed (Heracleum sphondylium) flowerhead, reacting to and catching a fly (Muscidae), Wiltshire, England, UK, June.
Two Dark green fritillary butterflies (Argynnis aglaja) and a Narrow-bordered five-spot burnet moth (Zygaena lonicerae) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland...
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Nick Upton / Two Dark green fritillary butterflies (Argynnis aglaja) and a Narrow-bordered five-spot burnet moth (Zygaena lonicerae) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland...
Six-spot burnet moth (Zygaena filipendulae), Marbled white butterflies (Melanargia galathea) and a Dark green fritillary (Argynnis aglaja) nectaring on a clump of Greater knapweed flowers (Centaurea s...
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Nick Upton / Six-spot burnet moth (Zygaena filipendulae), Marbled white butterflies (Melanargia galathea) and a Dark green fritillary (Argynnis aglaja) nectaring on a clump of Greater knapweed flowers (Centaurea s...
Male Thick-legged flower beetle (Oedemera nobilis), with a Pyramidal orchid (Anacamptis pyramidalis) pollinium attached to its head, nectaring on Common hogweed flowers (Heracleum sphondylium), Wiltsh...
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Nick Upton / Male Thick-legged flower beetle (Oedemera nobilis), with a Pyramidal orchid (Anacamptis pyramidalis) pollinium attached to its head, nectaring on Common hogweed flowers (Heracleum sphondylium), Wiltsh...
Marbled white butterflies (Melanargia galathea) nectaring on a clump of Greater knapweed flowers (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, England, UK, June.
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Nick Upton / Marbled white butterflies (Melanargia galathea) nectaring on a clump of Greater knapweed flowers (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, England, UK, June.
Marbled white butterflies (Melanargia galathea) nectaring on a clump of Greater knapweed flowers (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, England, UK, June.
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Nick Upton / Marbled white butterflies (Melanargia galathea) nectaring on a clump of Greater knapweed flowers (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, England, UK, June.
Feral horse (Equus caballus) walking on a remote sandy beach, near Arta, Mallorca, Spain, August.
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Nick Upton / Feral horse (Equus caballus) walking on a remote sandy beach, near Arta, Mallorca, Spain, August.
Forest caterpillar hunter (Calosoma sycophanta) feeding on a Gypsy moth caterpillar (Lymantria dispar) it has killed in a Holm oak tree, Bacu Goloritze ravine, Baunei, Sardinia, Italy, May.
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Nick Upton / Forest caterpillar hunter (Calosoma sycophanta) feeding on a Gypsy moth caterpillar (Lymantria dispar) it has killed in a Holm oak tree, Bacu Goloritze ravine, Baunei, Sardinia, Italy, May.
Sand-loving / Square-headed wasp (Tachysphex costae) feeding on Sea holly flowers (Eryngium maritimum) on a beach, Sardinia, Italy, June.
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Nick Upton / Sand-loving / Square-headed wasp (Tachysphex costae) feeding on Sea holly flowers (Eryngium maritimum) on a beach, Sardinia, Italy, June.
Captive reared juvenile White stork (Ciconia ciconia) in flight over the Knepp Estate soon after release, Sussex, UK, August 2019.
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Nick Upton / Captive reared juvenile White stork (Ciconia ciconia) in flight over the Knepp Estate soon after release, Sussex, UK, August 2019.
Captive reared juvenile White stork (Ciconia ciconia) with a GPS tracker on its back in flight over the Knepp Estate soon after release, Sussex, UK, August 2019.
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Nick Upton / Captive reared juvenile White stork (Ciconia ciconia) with a GPS tracker on its back in flight over the Knepp Estate soon after release, Sussex, UK, August 2019.
View towards Naranjo de Bulnes (Picu Urriellu) a distinctive 2519m limestone peak, Picos de Europa mountains, viewed from Arenas de Cabrales at sunset, Asturias, Spain, August 2016.
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Nick Upton / View towards Naranjo de Bulnes (Picu Urriellu) a distinctive 2519m limestone peak, Picos de Europa mountains, viewed from Arenas de Cabrales at sunset, Asturias, Spain, August 2016.
Mediterranean sea holly / Pyrenean eryngo (Eryngium bourgatii) flowering on montane pastureland, Covadonga, Picos de Europa, Asturias, Spain, August.
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Nick Upton / Mediterranean sea holly / Pyrenean eryngo (Eryngium bourgatii) flowering on montane pastureland, Covadonga, Picos de Europa, Asturias, Spain, August.
Pyrenean merendera / False meadow saffron (Merendera pyrenaica / Colchicum montanum) flowering on montane pastureland, Vegas de Sotres, Picos de Europa, Asturias, Spain, August.
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Nick Upton / Pyrenean merendera / False meadow saffron (Merendera pyrenaica / Colchicum montanum) flowering on montane pastureland, Vegas de Sotres, Picos de Europa, Asturias, Spain, August.
Cornish heath (Erica vagans) clump flowering on montane pastureland, above the Lakes of Covadonga, at 1300m, Picos de Europa, Asturias, Spain, August.
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Nick Upton / Cornish heath (Erica vagans) clump flowering on montane pastureland, above the Lakes of Covadonga, at 1300m, Picos de Europa, Asturias, Spain, August.
Pair of White storks (Ciconia ciconia) mating on their nest, Knepp Castle Estate, Sussex, England, UK, April. This is the first recorded instance of White storks nesting in the UK for several hundreds...
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Nick Upton / Pair of White storks (Ciconia ciconia) mating on their nest, Knepp Castle Estate, Sussex, England, UK, April. This is the first recorded instance of White storks nesting in the UK for several hundreds...
Pair of White storks (Ciconia ciconia) arranging nest material on their nest, Knepp Castle Estate, Sussex, England, UK, April. This is the first recorded instance of White storks nesting in the UK for...
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Nick Upton / Pair of White storks (Ciconia ciconia) arranging nest material on their nest, Knepp Castle Estate, Sussex, England, UK, April. This is the first recorded instance of White storks nesting in the UK for...
Pair of White storks (Ciconia ciconia) arranging nest material on their nest, Knepp Castle Estate, Sussex, England, UK, April. This is the first recorded instance of White storks nesting in the UK for...
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Nick Upton / Pair of White storks (Ciconia ciconia) arranging nest material on their nest, Knepp Castle Estate, Sussex, England, UK, April. This is the first recorded instance of White storks nesting in the UK for...
White stork (Ciconia ciconia) parent standing beside its three developing chicks. In captive breeding colony raising young birds for UK White Stork reintroduction project at the Knepp Estate. Cotswold...
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Nick Upton / White stork (Ciconia ciconia) parent standing beside its three developing chicks. In captive breeding colony raising young birds for UK White Stork reintroduction project at the Knepp Estate. Cotswold...
White stork (Ciconia ciconia) nest at captive breeding colony raising chicks for UK White Stork reintroduction project at the Knepp Estate. Cotswold Wildlife Park, Oxfordshire, UK, May 2019.
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Nick Upton / White stork (Ciconia ciconia) nest at captive breeding colony raising chicks for UK White Stork reintroduction project at the Knepp Estate. Cotswold Wildlife Park, Oxfordshire, UK, May 2019.
Richard Wardle feeding fish to recently hatched White stork (Ciconia ciconia) chicks in nest. In captive breeding colony raising young birds for UK White Stork reintroduction project at the Knepp Esta...
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Nick Upton / Richard Wardle feeding fish to recently hatched White stork (Ciconia ciconia) chicks in nest. In captive breeding colony raising young birds for UK White Stork reintroduction project at the Knepp Esta...
Recently hatched White stork (Ciconia ciconia) chicks begging for food in their nest. In captive breeding colony raising young birds for UK White Stork reintroduction project at the Knepp Estate. Cots...
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Nick Upton / Recently hatched White stork (Ciconia ciconia) chicks begging for food in their nest. In captive breeding colony raising young birds for UK White Stork reintroduction project at the Knepp Estate. Cots...
White stork (Ciconia ciconia) pair mating on their nest at sunset, Knepp estate, Sussex, UK, April 2019. This is the first recorded instance of White storks nesting in the UK for several hundreds of y...
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Nick Upton / White stork (Ciconia ciconia) pair mating on their nest at sunset, Knepp estate, Sussex, UK, April 2019. This is the first recorded instance of White storks nesting in the UK for several hundreds of y...
White stork (Ciconia ciconia) pair performing an up-down display with bill clattering on their nest in an Oak tree, Knepp estate, Sussex, UK, April 2019. This is the first recorded instance of White s...
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Nick Upton / White stork (Ciconia ciconia) pair performing an up-down display with bill clattering on their nest in an Oak tree, Knepp estate, Sussex, UK, April 2019. This is the first recorded instance of White s...
White stork (Ciconia ciconia) male landing with nest material and joining his mate on their nest in an Oak tree at sunset, Knepp estate, Sussex, UK, April 2019. This is the first recorded instance of...
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Nick Upton / White stork (Ciconia ciconia) male landing with nest material and joining his mate on their nest in an Oak tree at sunset, Knepp estate, Sussex, UK, April 2019. This is the first recorded instance of...
Red mason bees (Osmia bicornis) entering and leaving nest hole drilled into a log within an insect hotel, Gloucestershire, England, UK, April.
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Nick Upton / Red mason bees (Osmia bicornis) entering and leaving nest hole drilled into a log within an insect hotel, Gloucestershire, England, UK, April.
Ivy bee (Colletes hederae) mating ball, showing a mass of males clustered around a female recently emerged from her burrow, Wiltshire, England, UK, September.
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Nick Upton / Ivy bee (Colletes hederae) mating ball, showing a mass of males clustered around a female recently emerged from her burrow, Wiltshire, England, UK, September.
Two Red fox (Vulpes vulpes) cubs playing near the entrance to their den at dusk, Somerset, England, UK, May.
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Nick Upton / Two Red fox (Vulpes vulpes) cubs playing near the entrance to their den at dusk, Somerset, England, UK, May.
European eel (Anguilla anguilla) elver migrating upstream, using an eel pass on the side of a weir, Somerset Levels, England, UK, July.
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Nick Upton / European eel (Anguilla anguilla) elver migrating upstream, using an eel pass on the side of a weir, Somerset Levels, England, UK, July.
Male Eurasian beaver (Castor fiber) grazing at dusk, River Otter, Devon, England, UK, June.
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Nick Upton / Male Eurasian beaver (Castor fiber) grazing at dusk, River Otter, Devon, England, UK, June.
Tilt shot of a dam constructed by Eurasian beavers (Castor fiber) on a section of the River Otter, Devon, England, UK, June 2018.
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Nick Upton / Tilt shot of a dam constructed by Eurasian beavers (Castor fiber) on a section of the River Otter, Devon, England, UK, June 2018.
Female House spider (Tegenaria sp.) at the mouth of her tubular silk retreat in an old stone wall, Somerset, UK, September.
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Nick Upton / Female House spider (Tegenaria sp.) at the mouth of her tubular silk retreat in an old stone wall, Somerset, UK, September.
Running foliage spider / Spiny-legged sac spider (Liocranum rupicola), a nationally scarce species in the UK, hunting at night in an old stone wall, Wiltshire, UK, September.
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Nick Upton / Running foliage spider / Spiny-legged sac spider (Liocranum rupicola), a nationally scarce species in the UK, hunting at night in an old stone wall, Wiltshire, UK, September.
Leafcutter / Rose-cutter bee (Megachile willughbiella) lands at its nest in a Bamboo tube in an insect hotel near several nests she has completed and some Potter wasp (Ancistrocerus sp.) nests, Wiltsh...
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Nick Upton / Leafcutter / Rose-cutter bee (Megachile willughbiella) lands at its nest in a Bamboo tube in an insect hotel near several nests she has completed and some Potter wasp (Ancistrocerus sp.) nests, Wiltsh...
Common lace weaver / Lace-webbed spider (Amaurobius similis) male in its web in an old stone wall at night, Wiltshire, UK, September.
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Nick Upton / Common lace weaver / Lace-webbed spider (Amaurobius similis) male in its web in an old stone wall at night, Wiltshire, UK, September.
Common swifts (Apus apus) flying overhead, Wiltshire, UK, June.  Digital composite image.
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Nick Upton / Common swifts (Apus apus) flying overhead, Wiltshire, UK, June. Digital composite image.
Pine marten (Martes martes) male standing on a feeding table in a soft release cage after arrival from Scotland, during a reintroduction project by the Vincent Wildlife Trust, Cambrian Mountains, Wale...
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Nick Upton / Pine marten (Martes martes) male standing on a feeding table in a soft release cage after arrival from Scotland, during a reintroduction project by the Vincent Wildlife Trust, Cambrian Mountains, Wale...
Pine Marten (Martes martes) foraging at night in mixed coniferous and birch woodland in the area where live traps were set for a reintroduction project to Wales by the Vincent Wildlife Trust, Scottish...
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Nick Upton / Pine Marten (Martes martes) foraging at night in mixed coniferous and birch woodland in the area where live traps were set for a reintroduction project to Wales by the Vincent Wildlife Trust, Scottish...
Eurasian beaver (Castor fiber) adding a branch to its dam, Tayside, Scotland, UK, May. Filmed at night using a remote camera and infra red light.
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Nick Upton / Eurasian beaver (Castor fiber) adding a branch to its dam, Tayside, Scotland, UK, May. Filmed at night using a remote camera and infra red light.
Female Eurasian beaver (Castor fiber) and five kits feeding on a Willow (Salix) sapling, River Otter, Devon, England, UK, July. Part of Devon Wildlife Trust's Devon Beaver Trial.
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Nick Upton / Female Eurasian beaver (Castor fiber) and five kits feeding on a Willow (Salix) sapling, River Otter, Devon, England, UK, July. Part of Devon Wildlife Trust's Devon Beaver Trial.
Two Eurasian beaver kits (Castor fiber) feeding at night, born in the wild on the River Otter as part of the River Otter Beaver Trial, Devon, England, UK, August 2015.
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Nick Upton / Two Eurasian beaver kits (Castor fiber) feeding at night, born in the wild on the River Otter as part of the River Otter Beaver Trial, Devon, England, UK, August 2015.
Male Eurasian beaver (Castor fiber) at night, feeding on a branch in a pond, Devon Wildlife Trust's Devon Beaver Project, Devon, UK, April.
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Nick Upton / Male Eurasian beaver (Castor fiber) at night, feeding on a branch in a pond, Devon Wildlife Trust's Devon Beaver Project, Devon, UK, April.
Male Eurasian beaver (Castor fiber) at night, feeding on a branch in a pond, Devon Wildlife Trust's Devon Beaver Project, Devon, UK, April.
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Nick Upton / Male Eurasian beaver (Castor fiber) at night, feeding on a branch in a pond, Devon Wildlife Trust's Devon Beaver Project, Devon, UK, April.
Eurasian beaver (Castor fiber) at night, swimming, Devon Wildlife Trusts Devon Beaver Project enclosure, Devon, England, UK, March.
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Nick Upton / Eurasian beaver (Castor fiber) at night, swimming, Devon Wildlife Trusts Devon Beaver Project enclosure, Devon, England, UK, March.
Eurasian beaver (Castor fiber) at night, chewing bark and climbing along a felled tree, Devon Wildlife Trusts Devon Beaver Project enclosure, Devon, England, UK, March.
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Nick Upton / Eurasian beaver (Castor fiber) at night, chewing bark and climbing along a felled tree, Devon Wildlife Trusts Devon Beaver Project enclosure, Devon, England, UK, March.
Common starfish (Asterias rubens) moving in a rockpool, real time, Cornwall, England, UK, September.
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Nick Upton / Common starfish (Asterias rubens) moving in a rockpool, real time, Cornwall, England, UK, September.
Common prawns (Palaemon serratus) and a Rock goby (Gobius paganellus) feeding on a dead Montagu's crab (Xantho hydrophilus) in a rockpool, Cornwall, England, UK, September.
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Nick Upton / Common prawns (Palaemon serratus) and a Rock goby (Gobius paganellus) feeding on a dead Montagu's crab (Xantho hydrophilus) in a rockpool, Cornwall, England, UK, September.
Rock goby (Gobius paganellus) stealing food from a scavenging Common prawn (Palaemon serratus) in a rockpool, Cornwall, England, UK, September.
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Nick Upton / Rock goby (Gobius paganellus) stealing food from a scavenging Common prawn (Palaemon serratus) in a rockpool, Cornwall, England, UK, September.
Slow motion clip of a Meadow brown butterfly (Maniola jurtina) and a Gatekeeper butterfly (Pyronia tithonus) nectaring on Saw-wort flowers (Serratula tinctoria), with a Silver-washed fritillary (Argyn...
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Nick Upton / Slow motion clip of a Meadow brown butterfly (Maniola jurtina) and a Gatekeeper butterfly (Pyronia tithonus) nectaring on Saw-wort flowers (Serratula tinctoria), with a Silver-washed fritillary (Argyn...
Close up view of a Marbled white butterfly (Melanargia galathea) nectaring on a Field scabious (Knautia arvensis) flower, with another trying to court with it, Wiltshire, England, UK, July.
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Nick Upton / Close up view of a Marbled white butterfly (Melanargia galathea) nectaring on a Field scabious (Knautia arvensis) flower, with another trying to court with it, Wiltshire, England, UK, July.
Slow motion cllip of a Marbled white butterfly (Melanargia galathea) nectaring on a Creeping thistle (Cirsium arvense) flower, with another trying to court with it, Wiltshire, England, UK, July.
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Nick Upton / Slow motion cllip of a Marbled white butterfly (Melanargia galathea) nectaring on a Creeping thistle (Cirsium arvense) flower, with another trying to court with it, Wiltshire, England, UK, July.
Little egret (Egretta garzetta) struggling to swallow a large Pike (Esox lucius) it has just caught, Rutland Water, Rutland, England, UK, November.
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Nick Upton / Little egret (Egretta garzetta) struggling to swallow a large Pike (Esox lucius) it has just caught, Rutland Water, Rutland, England, UK, November.
Flock of Jackdaws (Corvus mondedula) taking off from their roost site, Gloucestershire, England, UK, December.
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Nick Upton / Flock of Jackdaws (Corvus mondedula) taking off from their roost site, Gloucestershire, England, UK, December.
Close-up of a Dark green fritillary (Argynnis aglaja) nectaring on a Greater knapweed (Centaurea scabiosa) flower and flying off, Wiltshire, England, UK, July.
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Nick Upton / Close-up of a Dark green fritillary (Argynnis aglaja) nectaring on a Greater knapweed (Centaurea scabiosa) flower and flying off, Wiltshire, England, UK, July.
Eurasian beaver (Castor fiber) feeding on grass amongst Meadow buttercups (Ranunculus acris) near a pond in a large enclosure, Devon, England, UK, June. Captive.
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Nick Upton / Eurasian beaver (Castor fiber) feeding on grass amongst Meadow buttercups (Ranunculus acris) near a pond in a large enclosure, Devon, England, UK, June. Captive.
Close up of a male Yellow dung fly (Scathophaga stercoraria) climbing up to the top of some sheep dung, Wiltshire, England, UK, March. Slow motion clip.
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Nick Upton / Close up of a male Yellow dung fly (Scathophaga stercoraria) climbing up to the top of some sheep dung, Wiltshire, England, UK, March. Slow motion clip.
Panning shot of partially flooded pastureland with large flocks of Wigeon (Anas penelope), Lapwings (Vanellus vanellus) and Golden plover (Pluvialis apricaria) in flight, Gloucestershire, England, UK,...
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Nick Upton / Panning shot of partially flooded pastureland with large flocks of Wigeon (Anas penelope), Lapwings (Vanellus vanellus) and Golden plover (Pluvialis apricaria) in flight, Gloucestershire, England, UK,...
Grey seal (Halichoerus grypus) pup aged one month, rubbing its head with its flipper, yawning and settling down to rest on a sandy beach, Norfolk, England, UK, January.
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Nick Upton / Grey seal (Halichoerus grypus) pup aged one month, rubbing its head with its flipper, yawning and settling down to rest on a sandy beach, Norfolk, England, UK, January.
Panning shot of a flock of Lapwings (Vanellus vanellus) landing on partially flooded pastureland near resting Black-tailed godwits (Limosa limosa) and grazing Wigeon (Anas penelope), Gloucestershire,...
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Nick Upton / Panning shot of a flock of Lapwings (Vanellus vanellus) landing on partially flooded pastureland near resting Black-tailed godwits (Limosa limosa) and grazing Wigeon (Anas penelope), Gloucestershire,...
Flock of Lapwings (Vanellus vanellus) in flight,  with Golden plover (Pluvialis apricaria) flying in the background, Gloucestershire, England, UK, January.
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Nick Upton / Flock of Lapwings (Vanellus vanellus) in flight, with Golden plover (Pluvialis apricaria) flying in the background, Gloucestershire, England, UK, January.
Common swift (Apus apus) feeding chick in a nest box, filmed using infrared light, Cambridge, England, UK, August.
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Nick Upton / Common swift (Apus apus) feeding chick in a nest box, filmed using infrared light, Cambridge, England, UK, August.
Young Roe deer (Capreolus capreolus) buck framed by ferns, Wiltshire garden, UK, October.
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Nick Upton / Young Roe deer (Capreolus capreolus) buck framed by ferns, Wiltshire garden, UK, October.
Brassy longhorn moth (Nemophora metallica) male animal standing on a Field scabious (Knautia arvensis) flower, the host plant for the larvae of this species, waiting for females to arrive, chalk grass...
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Nick Upton / Brassy longhorn moth (Nemophora metallica) male animal standing on a Field scabious (Knautia arvensis) flower, the host plant for the larvae of this species, waiting for females to arrive, chalk grass...
Young Edible crab (Cancer pagurus) and a Rock goby (Gobius paganellus) side by side in a rock pool, looking towards the camera, The Gower, Wales, UK, August.
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Nick Upton / Young Edible crab (Cancer pagurus) and a Rock goby (Gobius paganellus) side by side in a rock pool, looking towards the camera, The Gower, Wales, UK, August.
Cinnabar moth (Tyria jacobaeae) caterpillar feeding on Ragwort (Jacobaea vulgaris) in meadow. Wiltshire, England, UK. July.
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Nick Upton / Cinnabar moth (Tyria jacobaeae) caterpillar feeding on Ragwort (Jacobaea vulgaris) in meadow. Wiltshire, England, UK. July.
Goldenrod crab spider (Misumena vatia) female on Field scabious (Knautia arvensis) waiting for prey to land, legs raised, Wiltshire, England, UK. July.
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Nick Upton / Goldenrod crab spider (Misumena vatia) female on Field scabious (Knautia arvensis) waiting for prey to land, legs raised, Wiltshire, England, UK. July.
Eurasian beaver (Castor fiber) female swimming in river. River Otter tributary, Devon, England, UK. June.
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Nick Upton / Eurasian beaver (Castor fiber) female swimming in river. River Otter tributary, Devon, England, UK. June.
European badger (Meles meles) running across garden lawn at night, close to a house. Wiltshire, England, UK. May. Camera trap image.
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Nick Upton / European badger (Meles meles) running across garden lawn at night, close to a house. Wiltshire, England, UK. May. Camera trap image.
Grey squirrel (Sciurus carolinensis) feeding, sitting on garden lawn. Wiltshire, England, UK. May 2019. Camera trap image.
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Nick Upton / Grey squirrel (Sciurus carolinensis) feeding, sitting on garden lawn. Wiltshire, England, UK. May 2019. Camera trap image.
Common frog (Rana temporaria) frogspawn in pond created by damming of woodland stream by Eurasian beaver (Castor fiber). Beaver gnawed branch in background. Devon Beaver Project, Devon Wildlife Trust,...
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Nick Upton / Common frog (Rana temporaria) frogspawn in pond created by damming of woodland stream by Eurasian beaver (Castor fiber). Beaver gnawed branch in background. Devon Beaver Project, Devon Wildlife Trust,...
Common frog (Rana temporaria) frogspawn in pond created by damming of woodland stream by Eurasian beaver (Castor fiber). Beaver gnawed branch in background. Devon Beaver Project, Devon Wildlife Trust,...
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Nick Upton / Common frog (Rana temporaria) frogspawn in pond created by damming of woodland stream by Eurasian beaver (Castor fiber). Beaver gnawed branch in background. Devon Beaver Project, Devon Wildlife Trust,...
Male Common darter dragonfly (Sympetrum striolatum) sun basking on a fence post in early winter sunshine, Rutland Water, Rutland, UK, November.
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Nick Upton / Male Common darter dragonfly (Sympetrum striolatum) sun basking on a fence post in early winter sunshine, Rutland Water, Rutland, UK, November.
Male Common European Crane fly / Daddy long legs (Tipula paludosa) recently emerged and resting on Goose grass / Cleavers (Galium aparine) by a pond, Suffolk, UK, May.
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Nick Upton / Male Common European Crane fly / Daddy long legs (Tipula paludosa) recently emerged and resting on Goose grass / Cleavers (Galium aparine) by a pond, Suffolk, UK, May.
Caddis fly (Trichoptera) resting on riverbank vegetation, Wiltshire, UK, May.
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Nick Upton / Caddis fly (Trichoptera) resting on riverbank vegetation, Wiltshire, UK, May.
Silken cocoon of Narrow-bordered Five-spot Burnet moth (Zygaena lonicerae) attached to grass stem, chalk grassland meadow, Wiltshire, UK, May.
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Nick Upton / Silken cocoon of Narrow-bordered Five-spot Burnet moth (Zygaena lonicerae) attached to grass stem, chalk grassland meadow, Wiltshire, UK, May.
Brown mayfly (Ephemera vulgata) newly emerged, resting on riverside vegetation, River Avon, Lacock, Wiltshire, UK, May.
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Nick Upton / Brown mayfly (Ephemera vulgata) newly emerged, resting on riverside vegetation, River Avon, Lacock, Wiltshire, UK, May.
Brown mayfly (Ephemera vulgata) newly emerged on a riverside nettle leaf, River Avon, Lacock, Wiltshire, UK, May.
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Nick Upton / Brown mayfly (Ephemera vulgata) newly emerged on a riverside nettle leaf, River Avon, Lacock, Wiltshire, UK, May.
Bloody-nosed beetle (Timarcha tenebricosa) in a stand of its food plant Goose grass / Cleavers (Galium aparine) in a chalk grassland meadow, Wiltshire, UK, May.
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Nick Upton / Bloody-nosed beetle (Timarcha tenebricosa) in a stand of its food plant Goose grass / Cleavers (Galium aparine) in a chalk grassland meadow, Wiltshire, UK, May.
Male Black Snipe fly (Chrysopilus cristatus), standing on a Philadelphus leaf in a garden, Wiltshire, UK, May.
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Nick Upton / Male Black Snipe fly (Chrysopilus cristatus), standing on a Philadelphus leaf in a garden, Wiltshire, UK, May.
Black oil beetle (Meloe proscarabaeus) walking on a coast path, seeking a site to dig a burrow for its eggs, north Cornwall, UK, April.
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Nick Upton / Black oil beetle (Meloe proscarabaeus) walking on a coast path, seeking a site to dig a burrow for its eggs, north Cornwall, UK, April.
Black oil beetle (Meloe proscarabaeus) starting to excavate a nest burrow in a footpath on a chalk grassland hillside, the Ridgeway, Wiltshire, UK, April.
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Nick Upton / Black oil beetle (Meloe proscarabaeus) starting to excavate a nest burrow in a footpath on a chalk grassland hillside, the Ridgeway, Wiltshire, UK, April.
Black oil beetle (Meloe proscarabaeus) female excavating a nest burrow in coastal grassland, north Cornwall, UK, April.
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Nick Upton / Black oil beetle (Meloe proscarabaeus) female excavating a nest burrow in coastal grassland, north Cornwall, UK, April.
Beadlet anemone (Actinia equina) in a rock pool, Cornwall, UK, March.
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Nick Upton / Beadlet anemone (Actinia equina) in a rock pool, Cornwall, UK, March.
Alder fly (Sialis sp.) resting on riverside umbel stem, Wiltshire, UK, May.
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Nick Upton / Alder fly (Sialis sp.) resting on riverside umbel stem, Wiltshire, UK, May.
Velvet swimming crab (Necora puber) in a rock pool, hiding among Coralweed (Corallina officinalis), Cornwall, UK, March.
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Nick Upton / Velvet swimming crab (Necora puber) in a rock pool, hiding among Coralweed (Corallina officinalis), Cornwall, UK, March.
Strawberry anemone (Actinia fragracea) in a rock pool alongside Coralweed (Corallina officinalis), Cornwall, UK, March.
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Nick Upton / Strawberry anemone (Actinia fragracea) in a rock pool alongside Coralweed (Corallina officinalis), Cornwall, UK, March.
Spotted longhorn beetle (Rutpela maculata = Strangalia maculata) takes off after feeding on a Common spotted orchid flower (Dactylorhiza fuchsii) in a woodland ride, Lower Woods, Gloucestershire, UK,...
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Nick Upton / Spotted longhorn beetle (Rutpela maculata = Strangalia maculata) takes off after feeding on a Common spotted orchid flower (Dactylorhiza fuchsii) in a woodland ride, Lower Woods, Gloucestershire, UK,...
Common shore crab (Carcinus maenas) hiding among Coralweed (Corallina officinalis) in a rock pool, Cornwall, UK, March.
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Nick Upton / Common shore crab (Carcinus maenas) hiding among Coralweed (Corallina officinalis) in a rock pool, Cornwall, UK, March.
Young female Nursery web spider (Pisaura mirabilis) sun basking on a leaf on a late winter day, Wiltshire garden, UK, February.
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Nick Upton / Young female Nursery web spider (Pisaura mirabilis) sun basking on a leaf on a late winter day, Wiltshire garden, UK, February.
Dense bed of Common mussels (Mytilus edulis) exposed at low tide on rocks fringing a sandy beach, Godrevy, Cornwall, UK, April.
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Nick Upton / Dense bed of Common mussels (Mytilus edulis) exposed at low tide on rocks fringing a sandy beach, Godrevy, Cornwall, UK, April.
Meadow plant bug (Leptopterna dolabrata) female laying an egg on a grass blade in a meadow, Wiltshire, UK, June.
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Nick Upton / Meadow plant bug (Leptopterna dolabrata) female laying an egg on a grass blade in a meadow, Wiltshire, UK, June.
Meadow brown butterfly (Maniola jurtina) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, UK, June.
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Nick Upton / Meadow brown butterfly (Maniola jurtina) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, UK, June.
Marbled white butterfly (Melanargia galathea) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, UK, June.
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Nick Upton / Marbled white butterfly (Melanargia galathea) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, UK, June.
Female Kestrel (Falco tinnunculus) soliciting male to mate with her by crouching down on a cliff ledge near their nest site, Cornwall, UK, April.
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Nick Upton / Female Kestrel (Falco tinnunculus) soliciting male to mate with her by crouching down on a cliff ledge near their nest site, Cornwall, UK, April.
Kestrel (Falco tinnunculus) female perched on a cliff ledge while eating a Common shrew (Sorex araneus) brought by her mate as a courtship gift and held in her talons, Cornwall, UK, April.
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Nick Upton / Kestrel (Falco tinnunculus) female perched on a cliff ledge while eating a Common shrew (Sorex araneus) brought by her mate as a courtship gift and held in her talons, Cornwall, UK, April.