Floris van Breugel / Sand dunes reflecting pink and orange glowing colours from the sky at sunset. Death Valley National Park, California, USA. November 2009. Highly Commended in the Veolia Environnement Wildlife Photoga...

George Sanker / Ellis River high in White Mountain National Forest. New Hampshire, October 2012.

George Sanker / American Buffalo / Bison (Bison bison) grazing in open plains. Yellowstone National Park, Wyoming, October.

Gavin Hellier / Wenkchemna Peaks or Ten Peaks rising over Moraine lake in the snow, near Lake Louise, Banff National Park, Alberta, Canada, 2007

Gavin Hellier / Wenkchemna Peaks or Ten Peaks rising over Moraine lake in the snow, near Lake Louise, Banff National Park, Alberta, Canada, 2007

Gavin Hellier / Peyto Lake, coloured by glacial silt, Banff-Jasper National Parks, Alberta, Canada, 2007

Jeff Vanuga / Aerial view of Grand Prismatic Springs Midway Geyser Basin. Yellowstone National Park, Wyoming, July 2011.

Wild Wonders of Europe / Widstrand / Sunrise over North Velebit National Park, Velebit Nature Park, Rewilding Europe area, Velebit mountains, Croatia June 2012

Tim Fitzharris / Ruby Beach, Olympic National Park, Washington

Tim Fitzharris / Mount Rundle and Johnson Lake, Banff National Park, Alberta, Canada

Sumio Harada / Black Bear (Ursus americanus) mother and cub near tree den, Glacier National Park, Montana

Kevin Schafer / Larch (Larix sp) trees in autumn below Mount Assiniboine with Sunburst Lake, Mount Assiniboine Provincial Park, British Columbia, Canada

Ingo Arndt / Grizzly Bear (Ursus arctos horribilis) front and rear tracks in mud, Lake Clark National Park, Alaska

Ingo Arndt / Grizzly Bear (Ursus arctos horribilis) chasing salmon, Lake Clark National Park, Alaska

Ingo Arndt / Grizzly Bear (Ursus arctos horribilis) cub running through shallow water, Lake Clark National Park, Alaska

Ingo Arndt / Grizzly Bear (Ursus arctos horribilis) mother with cub, Lake Clark National Park, Alaska

Mark Carwardine / Domestic dog wearing muzzle, trained to track the endangered North Island Kiwi, Trounson Kauri Park, North Island, New Zealand, January 2009

Floris van Breugel / Autumn colors along the shores of Pharaoh Lake in Adirondacks park, New York, USA. October

Floris van Breugel / Winter snowstorm with frost covered trees , Hurricane Ridge, Olympic National Park, Washington, USA, February 2012.

Floris van Breugel / Sub-alpine larches (Latrix sp.) in late autumn in North Cascades National Park, Washington, USA. October.

Floris van Breugel / A skier glides past an erupting geyser in the Old Faithful geyser basin in Yellowstone National Park. The cold temperatures in the winter cause the geysers to release more steam than usual making the...

Floris van Breugel / Natural iridescent oil film from decaying leaves at the surface of a pool, with a cottonwood leaf floating. Escalante National Monument, Utah. November.

Floris van Breugel / Hoar frost coats the edges of ferns in a clearing in Washington's Olympic National Park, USA. December.

Jack Dykinga / Bark patterns of Aspen tree (Populus tremula) in front of a Ponderosa pine (Pinus ponderosa) trunk, Kaibab National Forest, Arizona, USA

Jack Dykinga / Sunflower flower (Helianthus petiolaris) at Sunset Crater National Monument, Arizona

Oriol Alamany / Angel Glacier with morraine, Jasper National Park, Rocky Mountains, Alberta, Canada

Oriol Alamany / Nigel Creek with conifer forest at dusk, Banff National Park, Rocky Mountains, Alberta, Canada

Steven Kazlowski / Killer whale (Orcinus orca) large pod with one male approaching, in Kenai Fjords National Park, Chiswell Islands National Marine Sanctuary, South Central Alaska, USA

Steven Kazlowski / Moose (Alces alces) bull adult swims across a lake in Denali National Park, Interior of Alaska, summer, USA

Steven Kazlowski / Grizzly bear (Ursus arctos horribilis) young rare blonde (white) bear plays with a young dark coloured bear along the coast of Katmai National Park, Alaskan peninsula, USA

Steven Kazlowski / Kayaker in front of Bear Glacier, Kenai Fjords National Park, Alaska, USA. No release available.

Steven Kazlowski / Dall sheep (Ovis dalli) solitary animal at sunset with a storm moving across the sky, Denali National Park, interior Alaska, USA

Loic Poidevin / Grizzly bear (Ursus arctos horribilis) attempting to catch salmon leaping up rapids, Katmai National Park, Alaska, USA

Loic Poidevin / Salmon landing on head of Grizzly bear (Ursus arctos horribilis) as it is leaping up rapids, Katmai National Park, Alaska, USA

Loic Poidevin / Grizzly bear (Ursus arctos horribilis) salmon fishing in river rapids, Katmai National Park, Alaska, USA

Loic Poidevin / Grizzly bear (Ursus arctos horribilis) salmon fishing in river rapids, Katmai National Park, Alaska, USA

Jack Dykinga / Matkatamiba Canyon with its Muave limestone scoured by erosion. Grand Canyon National Park, Arizona. May 2012.

Jack Dykinga / Shore Acres State Park with sandstone bluffs being pounded by tidal shifting surf, creating waterfalls over concretions in the stone. Oregon

Mark Taylor / Forest fire at night with silhouetted trees, Yellowstone National Park, Wyoming, USA

Paul D Stewart / Aerial view of Grand Prismatic thermal spring in Yellowstone National Park, USA. The bright colours are natural, coming from thermophile bacteria in the water. People can be seen walking on the raised...

Juan Carlos Munoz / Great Egret (Ardea alba) in full breeding plumage. Everglades National Park, Florida, USA, February.

Juan Carlos Munoz / Great Egrets (Ardea alba) territorial fight above nest colony, Everglades National Park, Florida, USA, February.

Juan Carlos Munoz / Wood Stork / American Wood Ibis (Mycteria americana) in flight. Everglades National Park, Florida, USA, February.

Juan Carlos Munoz / Great Egrets (Ardea alba) territorial dispute above nest colony, Everglades National Park, Florida, USA, February.

Juan Carlos Munoz / Black-crowned Night Heron (Nycticorax nycticorax) portrait. Everglades National Park, Florida, USA, February.

Juan Carlos Munoz / Roseate Spoonbill (Platalea ajaja) rear view wings open, in mangrove canopy. Everglades National Park, Florida, USA, February.

Juan Carlos Munoz / Roseate Spoonbill (Platalea ajaja) in flight. Everglades National Park, Florida, USA, February.

Juan Carlos Munoz / Snowy Egret (Egretta thula) portrait. Everglades National Park, Florida, USA, February.

Juan Carlos Munoz / Snowy Egret (Egretta thula) talon. Everglades National Park, Florida, USA, February.

Juan Carlos Munoz / Osprey (Pandion haliaetus) in flight with caught fish. Everglades National Park, Florida, USA, August.

Juan Carlos Munoz / Double Breasted Cormorant (Phalacrocorax auritus) resting beak in plumage. Everglades National Park, Florida, USA, February.

Juan Carlos Munoz / Snowy Egret (Egretta thula) reflected in water. Everglades National Park, Florida, USA, February.

Juan Carlos Munoz / Aerial view over subtropical mangrove wetlands of the Everglades National Park. Florida, USA, February 2012.

Juan Carlos Munoz / Aerial view over subtropical mangrove wetlands of the Everglades National Park. Florida, USA, February 2012.

Juan Carlos Munoz / Aerial view over subtropical mangrove wetlands of the Everglades National Park. Florida, USA, February 2012.

Juan Carlos Munoz / Aerial view over subtropical mangrove wetlands of the Everglades National Park. Florida, USA, February 2012.

Steven Kazlowski / Moose (Alces alces) newborn calf kneels to feed on vegetation. Kincaid Park, Anchorage, south-central Alaska, May.

Steven Kazlowski / Killer WHale / Orca (Orcinus orca) bull and cow in Resurrection Bay, Kenai Fjords National Park, outside Seward, Alaska, May.

George Sanker / American darter / Anhinga (Anhinga anhinga) with caught fish prey, Everglades National Park, Florida, USA, March

George Sanker / Purple gallinule (Porphyrio martinica) moving across waterlily leaves, Everglades National Park, Florida, USA, April

George Sanker / Purple gallinule (Porphyrio martinica) moving across waterlily covered surface, Everglades National Park, Florida, USA, March

George Sanker / Green heron (Butorides virescens) hunting for fish, camouflaged against tree roots, Everglades National Park, Florida, USA, March

George Sanker / Black bear (Ursus americanus) young cub portrait, Yellowstone National Park, Wyoming, USA

Andy Rouse / Dingo (Canis lupus dingo) Kakadu National Park, North West Territories, Australia

Jack Dykinga / Sand storm cutting across the Mesquite Sand Dunes in Death Valley National Park, California, USA 2010

Jack Dykinga / Cheyenne River Sioux Tribal Park with severe storm clouds at sunset amid vast grassland, South Dakota, USA

Jack Dykinga / North Rim at Cape Royal with sunset storm light streaming in rays, backlighting the canyon ridges, Grand Canyon National Park, Arizona, USA

Bertie Gregory / Raccoon (Procyon lotor) standing up investigating camera, portrait, Stanley park, Vancouver, British Columbia, Cananda, September.

Bertie Gregory / Raccoon (Procyon lotor) two, portraits showing hands and claws, Stanley park, Vancouver, British Columbia, Cananda, September.

Juan Carlos Munoz / Aerial view of Everglades National Park, Florida, USA, February 2012

Juan Carlos Munoz / Aerial view of Everglades National Park, Florida, USA, February 2012

Danny Green / Bison (Bison bison) seen through blizzard snow. Yellowstone National Park, USA, February.

Danny Green / Bison (Bison bison) in snow. Yellowstone National Park, USA, February.

Danny Green / Bison (Bison bison) in snowstorm. Yellowstone National Park, USA, February.

Danny Green / Bison (Bison bison) in snowstorm. Yellowstone National Park, USA, February.

Danny Green / Bison (Bison bison) in snowstorm. Yellowstone National Park, USA, February.

Konstantin Mikhailov / Lake Ann surrounded by Douglas firs (Pseudotsuga menziesii) Silver firs (Abies alba) and other conifers in the area of Rainy Pass of North Cascades National Park, North Cascades National Park, Cascade...

Konstantin Mikhailov / Western red cedar tree (Thuja plicata) trunks with moss and fungi, inland temperate rainforest of Olympic National Park, Washington, USA, Western North America, January.

Claudio Contreras / Tres Marias Rabbit (Sylvilagus graysoni) in profile. Maria Magdalena Island, Islas Marias Biosphere Reserve, Sea of Cortez (Gulf of California), Mexico, June. Endangered.

Claudio Contreras / Tres Marias Rabbit (Sylvilagus graysoni). Maria Madre Island, Islas Marias Biosphere Reserve, Sea of Cortez (Gulf of California), Mexico, July. Endangered.

Claudio Contreras / Mexican / Brown Vine Snake (Oxybelis aeneus) with mouth open. Maria Madre Island, Islas Marias Biosphere Reserve, Sea of Cortez (Gulf of California), Mexico, June.

Dave Watts / Southern cassowary (Casuarius casuarius) adult male with chicks, World Heritage National Park rainforest of the Wet Tropics, north Queensland, Australia

Kirkendall-Spring / Sunset over West and Middle Anacapa Islands, view from Inspiration Point on East Anacapa Island, Channel Islands National Park, California, USA, April 2011.

George Sanker / Bison (Bison bison) two males head to head, Yellowstone NP, Wyoming, USA

George Sanker / Black Bear (Ursus americanus) cub balancing on branch in tree, Yellowstone NP, Wyoming, USA

George Sanker / Coyote (Canis latrans) standing on rocks at water's edge, sniffing, Yellowstone NP, Wyoming, USA, October

George Sanker / Grizzly bear (Ursus arctos horribilis) walking through long grass, Yellowstone NP, Wyoming, USA, October

George Sanker / American beaver (Castor canadensis) swimming, Grand Teton NP, Wyoming, USA, October

George Sanker / Least Chipmunk (Tamius minimus) feeding on the seeds of a thistle bush during a snowstorm, Grand Teton NP, Wyoming, USA, October

Gerrit Vyn / Dall sheep (Ovis dalli) rams head butting, one behaviour by which males establish dominance during the Autumn mating season. Denali National Park, Alaska, USA

Gerrit Vyn / Dall sheep (Ovis dalli) yearling lamb running across alpine tundra, Denali National Park, Alaska, USA

George Sanker / Grizzly bear (Ursus arctos horribilis) walking, Yellowstone National Park, Wyoming, USA, October

George Sanker / Long-tailed weasel (Mustela frenata) standing on hind legs, Yellowstone National Park, Wyoming, USA, June

George Sanker / Two Least Chipmunks (Tamias minimus) sitting on tree stump feeding, Yellowstone National Park, Wyoming, USA, October, crop of 1373882

George Sanker / Pond in forest, Baxter State Park, Maine, USA, October 2007

George Sanker / Newborn Elk (Cervus canadensis) calf looks up to its mother, Yellowstone National Park, USA, June