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Female Capybara (Hydrochoerus hydrochaeris) swimming in line with young after escaping a Jaguar attack (Panthera onca palustris) in a lagoon off the Paraguay River. Taiama Ecological Reserve, Pantanal...
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Nick Garbutt / Female Capybara (Hydrochoerus hydrochaeris) swimming in line with young after escaping a Jaguar attack (Panthera onca palustris) in a lagoon off the Paraguay River. Taiama Ecological Reserve, Pantanal...
Yacare Caiman (Caiman yacare) gaping to regulate its body temperature, with attendant hoverflies, the Paraguay River, Taiama Reserve, western Pantanal, Brazil, South America.
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Nick Garbutt / Yacare Caiman (Caiman yacare) gaping to regulate its body temperature, with attendant hoverflies, the Paraguay River, Taiama Reserve, western Pantanal, Brazil, South America.
Yellow Anaconda (Eunectes notaeus) in vegetation at the edge of the Paraguay River, Taiama Reserve, western Pantanal, Brazil, South America.
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Nick Garbutt / Yellow Anaconda (Eunectes notaeus) in vegetation at the edge of the Paraguay River, Taiama Reserve, western Pantanal, Brazil, South America.
River stingray (Potamotrygon sp.), Paraguay River, Pantanal, Brazil
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Franco Banfi / River stingray (Potamotrygon sp.), Paraguay River, Pantanal, Brazil
Streaked prochilod (Prochilodus lineatus), species native to the Parana-Paraguay and Paraiba do Sul river basins in South America. Recanto Ecologico, Rio da Prata, Bonito, Mato Grosso do Sul, Brazil
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Franco Banfi / Streaked prochilod (Prochilodus lineatus), species native to the Parana-Paraguay and Paraiba do Sul river basins in South America. Recanto Ecologico, Rio da Prata, Bonito, Mato Grosso do Sul, Brazil
Red-bellied piranha (Pygocentrus nattereri),  on a secondary river from the Paraguay River, Pantanal, Brazil
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Franco Banfi / Red-bellied piranha (Pygocentrus nattereri), on a secondary river from the Paraguay River, Pantanal, Brazil
Red-bellied piranha (Pygocentrus nattereri),  on a secondary river from the Paraguay River, Pantanal, Brazil
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Franco Banfi / Red-bellied piranha (Pygocentrus nattereri), on a secondary river from the Paraguay River, Pantanal, Brazil
Water hyacinth, (Eichhornia sp.), from below the surface, Paraguay River, Pantanal, Brazil
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Franco Banfi / Water hyacinth, (Eichhornia sp.), from below the surface, Paraguay River, Pantanal, Brazil
River stingray (Potamotrygon sp.), Paraguay River, Pantanal, Brazil
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Franco Banfi / River stingray (Potamotrygon sp.), Paraguay River, Pantanal, Brazil
River stingray (Potamotrygon sp.), Paraguay River, Pantanal, Brazil
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Franco Banfi / River stingray (Potamotrygon sp.), Paraguay River, Pantanal, Brazil
Red-bellied piranha (Pygocentrus nattereri ), at night, in tributary of Paraguay River, Pantanal, Brazil
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Franco Banfi / Red-bellied piranha (Pygocentrus nattereri ), at night, in tributary of Paraguay River, Pantanal, Brazil
Boat loaded to travel the Paraguay river for exploration, Pantanal, Brazil
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Franco Banfi / Boat loaded to travel the Paraguay river for exploration, Pantanal, Brazil
Snorkelling the tributary of the Paraguay River, Pantanal, Brazil
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Franco Banfi / Snorkelling the tributary of the Paraguay River, Pantanal, Brazil
Detail of the leg of a Yacare caiman (Caiman yacare), Paraguay river, Pantanal wetlands, Mato Grosso, Brazil
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Franco Banfi / Detail of the leg of a Yacare caiman (Caiman yacare), Paraguay river, Pantanal wetlands, Mato Grosso, Brazil
Yacare caiman (Caiman yacare), Paraguay river, Pantanal wetlands, Mato Grosso, Brazil
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Franco Banfi / Yacare caiman (Caiman yacare), Paraguay river, Pantanal wetlands, Mato Grosso, Brazil
Yacare caiman (Caiman yacare), Paraguay river, Pantanal wetlands, Mato Grosso, Brazil
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Franco Banfi / Yacare caiman (Caiman yacare), Paraguay river, Pantanal wetlands, Mato Grosso, Brazil
Yacare caiman (Caiman yacare) Paraguay river, Pantanal wetlands, Mato Grosso, Brazil
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Franco Banfi / Yacare caiman (Caiman yacare) Paraguay river, Pantanal wetlands, Mato Grosso, Brazil
Red-bellied piranha (Pygocentrus nattereri ), in tributary at night, Paraguay River, Pantanal, Brazil
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Franco Banfi / Red-bellied piranha (Pygocentrus nattereri ), in tributary at night, Paraguay River, Pantanal, Brazil
Trees reflected in the a tributary river connected to Paraguay River, Pantanal, Brazil
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Franco Banfi / Trees reflected in the a tributary river connected to Paraguay River, Pantanal, Brazil
Yacare caiman (Caiman yacare),basking in the sun, Paraguay river, Pantanal wetlands, Mato Grosso, Brazil
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Franco Banfi / Yacare caiman (Caiman yacare),basking in the sun, Paraguay river, Pantanal wetlands, Mato Grosso, Brazil
Yacare caiman (Caiman yacare), peering out of water, Paraguay river, Pantanal wetlands, Mato Grosso, Brazil
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Franco Banfi / Yacare caiman (Caiman yacare), peering out of water, Paraguay river, Pantanal wetlands, Mato Grosso, Brazil
View finder of camera showing Yacare caiman (Caiman yacare), Paraguay river, Pantanal wetlands, Mato Grosso, Brazil
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Franco Banfi / View finder of camera showing Yacare caiman (Caiman yacare), Paraguay river, Pantanal wetlands, Mato Grosso, Brazil
Jaguar (Panthera onca) female swimming across the Paraguay River. Near Taiama Reserve, Western Pantanal, Mato Grosso, Brazil.
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Nick Garbutt / Jaguar (Panthera onca) female swimming across the Paraguay River. Near Taiama Reserve, Western Pantanal, Mato Grosso, Brazil.
Jaguar (Panthera onca) female swimming across the Paraguay River. Near Taiama Reserve, Western Pantanal, Mato Grosso, Brazil.
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Nick Garbutt / Jaguar (Panthera onca) female swimming across the Paraguay River. Near Taiama Reserve, Western Pantanal, Mato Grosso, Brazil.
A small flock of Roseatte spoonbills (Ajaia ajaja) in flight. Paraguay River, Taiama Reserve, Western Pantanal, Mato Grosso, Brazil,
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Nick Garbutt / A small flock of Roseatte spoonbills (Ajaia ajaja) in flight. Paraguay River, Taiama Reserve, Western Pantanal, Mato Grosso, Brazil,
Jaguar (Panthera onca) female swimming across the Paraguay River. Near Taiama Reserve, Western Pantanal, Mato Grosso, Brazil.
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Nick Garbutt / Jaguar (Panthera onca) female swimming across the Paraguay River. Near Taiama Reserve, Western Pantanal, Mato Grosso, Brazil.
Yellow king piranha (Pygocentrus natteri), Paraguay River, Pantanal, Brazil
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Franco Banfi / Yellow king piranha (Pygocentrus natteri), Paraguay River, Pantanal, Brazil
Camera viewfinder with Yacare caiman (Caiman yacare) in frame, Paraguay river, Pantanal wetlands, Mato Grosso, Brazil
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Franco Banfi / Camera viewfinder with Yacare caiman (Caiman yacare) in frame, Paraguay river, Pantanal wetlands, Mato Grosso, Brazil
Aerial view of Corumba, at end of the dry season, with the Rio Paraguay or Paraguay river, Brazil. November 2017. Photographed for The Freshwater Project
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Michel Roggo / Aerial view of Corumba, at end of the dry season, with the Rio Paraguay or Paraguay river, Brazil. November 2017. Photographed for The Freshwater Project
Aerial view of the Pantanal, end of the dry season. Near the Rio Paraguay or Paraguay river, Brazil. November 2017. Photographed for The Freshwater Project
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Michel Roggo / Aerial view of the Pantanal, end of the dry season. Near the Rio Paraguay or Paraguay river, Brazil. November 2017. Photographed for The Freshwater Project
Aerial view of the Pantanal around the Rio Paraguay or Paraguay River, at the end of the dry season, Brazil. November. Photographed for The Freshwater Project
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Michel Roggo / Aerial view of the Pantanal around the Rio Paraguay or Paraguay River, at the end of the dry season, Brazil. November. Photographed for The Freshwater Project
Aerial view of the Pantanal, end of the dry season. Near the  area of the Rio Paraguay or Paraguay river, north of Corumba, Brazil. November. Photographed for the Freshwater Project.
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Michel Roggo / Aerial view of the Pantanal, end of the dry season. Near the area of the Rio Paraguay or Paraguay river, north of Corumba, Brazil. November. Photographed for the Freshwater Project.
Aerial view of the Pantanal, at the end of the dry season, near the Paraguay River, Brazil. November 2016. Photographed for The Freshwater Project
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Michel Roggo / Aerial view of the Pantanal, at the end of the dry season, near the Paraguay River, Brazil. November 2016. Photographed for The Freshwater Project
River stingray (Potamotrygon sp.) half hidden in sand, Paraguay River, Pantanal, Brazil
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Franco Banfi / River stingray (Potamotrygon sp.) half hidden in sand, Paraguay River, Pantanal, Brazil
River stingray (Potamotrygon sp.), Paraguay River, Pantanal, Brazil
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Franco Banfi / River stingray (Potamotrygon sp.), Paraguay River, Pantanal, Brazil
River stingray (Potamotrygon sp.), Paraguay River, Pantanal, Brazil
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Franco Banfi / River stingray (Potamotrygon sp.), Paraguay River, Pantanal, Brazil
River stingray (Potamotrygon sp.) camouflaged in sand, Paraguay River, Pantanal, Brazil
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Franco Banfi / River stingray (Potamotrygon sp.) camouflaged in sand, Paraguay River, Pantanal, Brazil
Giant Otter (Pteronura brasiliensis) holding onto a branch in a lagoon off the Paraguay River, Taiama Reserve, western Pantanal, Brazil, South America.
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Nick Garbutt / Giant Otter (Pteronura brasiliensis) holding onto a branch in a lagoon off the Paraguay River, Taiama Reserve, western Pantanal, Brazil, South America.
Female Capybara (Hydrochoerus hydrochaeris) swimming with young and alarm calling after escaping a Jaguar attack (Panthera onca palustris) in a lagoon off the Paraguay River, Taiama Ecological Reserve...
01493078
Nick Garbutt / Female Capybara (Hydrochoerus hydrochaeris) swimming with young and alarm calling after escaping a Jaguar attack (Panthera onca palustris) in a lagoon off the Paraguay River, Taiama Ecological Reserve...
Jabiru Stork (Jabiru mycteria) catching a fish (Pacu) on the edge of the Paraguay River. Taiama Ecological Reserve, Pantanal, Brazil, South America.
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Nick Garbutt / Jabiru Stork (Jabiru mycteria) catching a fish (Pacu) on the edge of the Paraguay River. Taiama Ecological Reserve, Pantanal, Brazil, South America.
Jabiru Stork (Jabiru mycteria) catching a fish (Pacu) on the edge of the Paraguay River. Taiama Ecological Reserve, Pantanal, Brazil, South America.
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Nick Garbutt / Jabiru Stork (Jabiru mycteria) catching a fish (Pacu) on the edge of the Paraguay River. Taiama Ecological Reserve, Pantanal, Brazil, South America.
Black-capped Donacobius (Donacobius atricapilla) singing from floating Water Hyacinth (Eichhornia sp), Taiama Ecological Reserve, Paraguay River, Pantanal, Brasil.
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Nick Garbutt / Black-capped Donacobius (Donacobius atricapilla) singing from floating Water Hyacinth (Eichhornia sp), Taiama Ecological Reserve, Paraguay River, Pantanal, Brasil.
Jabiru Stork (Jabiru mycteria) catching a fish (Pacu) on the edge of the Paraguay River, Taiama Ecological Reserve, Pantanal, Brazil, South America.
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Nick Garbutt / Jabiru Stork (Jabiru mycteria) catching a fish (Pacu) on the edge of the Paraguay River, Taiama Ecological Reserve, Pantanal, Brazil, South America.
Jacare Caiman (Caiman yacare), Pantanal, Mato Grosso, Brazil
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Pete Oxford / Jacare Caiman (Caiman yacare), Pantanal, Mato Grosso, Brazil
'Novateiro' Tree (Triplaris brasiliana) flowering on the shore of Paraguay River at the Pantanal of Mato Grosso, Taiama Ecological Station, Mato Grosso State, Western Brazil
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Luiz Claudio Marigo / 'Novateiro' Tree (Triplaris brasiliana) flowering on the shore of Paraguay River at the Pantanal of Mato Grosso, Taiama Ecological Station, Mato Grosso State, Western Brazil
Paraguay Caiman (Caiman yacare) feeding on fish, on banks of Piquiri River, Pantanal of Mato Grosso, Mato Grosso State, Western Brazil.
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Luiz Claudio Marigo / Paraguay Caiman (Caiman yacare) feeding on fish, on banks of Piquiri River, Pantanal of Mato Grosso, Mato Grosso State, Western Brazil.
Paraguay Caiman (Caiman yacare) feeding on fish, in Piquiri River, Pantanal of Mato Grosso, Mato Grosso State, Western Brazil.
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Luiz Claudio Marigo / Paraguay Caiman (Caiman yacare) feeding on fish, in Piquiri River, Pantanal of Mato Grosso, Mato Grosso State, Western Brazil.
Paraguay Caiman (Caiman yacare) feedin on fish, in Piquiri River, Pantanal of Mato Grosso, Mato Grosso State, Western Brazil.
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Luiz Claudio Marigo / Paraguay Caiman (Caiman yacare) feedin on fish, in Piquiri River, Pantanal of Mato Grosso, Mato Grosso State, Western Brazil.
Aerial view of the Pantanal, at the end of the dry season, area of the Rio Paraguay or Paraguay river, north of Corumba, Brazil, November 2012. Taken for the Freshwater Project.
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Michel Roggo / Aerial view of the Pantanal, at the end of the dry season, area of the Rio Paraguay or Paraguay river, north of Corumba, Brazil, November 2012. Taken for the Freshwater Project.
Puiva Trees (Tabebuia impetiginosa) in full bloom. Near the banks of the Paraguay River, Taiama Reserve, Mato Grosso, Western Pantanal, Brazil.
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Nick Garbutt / Puiva Trees (Tabebuia impetiginosa) in full bloom. Near the banks of the Paraguay River, Taiama Reserve, Mato Grosso, Western Pantanal, Brazil.
Jacare Caiman (Caiman yacare) emerging from river, Pantanal, Brazil
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Luciano Candisani / Jacare Caiman (Caiman yacare) emerging from river, Pantanal, Brazil
Jacare Caiman (Caiman yacare) and rainbow during dry season, Pantanal, Brazil
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Luciano Candisani / Jacare Caiman (Caiman yacare) and rainbow during dry season, Pantanal, Brazil
Aquatic plants, Rio Paraguay, Pantanal, Brazil
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Luciano Candisani / Aquatic plants, Rio Paraguay, Pantanal, Brazil
Jacare Caiman (Caiman yacare) gathering at water's edge during dry season, Pantanal, Brazil
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Luciano Candisani / Jacare Caiman (Caiman yacare) gathering at water's edge during dry season, Pantanal, Brazil
Group of Red-and-Green Macaws  (Ara chloropterus) in flight over forest canopy. Taiama Ecological Reserve, Paraguay River, Pantanal, Brazil.
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Nick Garbutt / Group of Red-and-Green Macaws (Ara chloropterus) in flight over forest canopy. Taiama Ecological Reserve, Paraguay River, Pantanal, Brazil.
Hyacinth Macaws (Anodorhynchus hyacinthinus) in flight. Taiama Ecological Reserve, Paraguay River, Pantanal, Brazil.
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Nick Garbutt / Hyacinth Macaws (Anodorhynchus hyacinthinus) in flight. Taiama Ecological Reserve, Paraguay River, Pantanal, Brazil.
Southern Screamer or Crested Screamer (Chauna torquata) in flight and landing on river-side tree. Taiama Ecological Reserve, Paraguay River, western Pantanal, Brazil. Digital composite
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Nick Garbutt / Southern Screamer or Crested Screamer (Chauna torquata) in flight and landing on river-side tree. Taiama Ecological Reserve, Paraguay River, western Pantanal, Brazil. Digital composite
Jacare Caiman (Caiman yacare) fishing in flowing river, Pantanal, Brazil
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Luciano Candisani / Jacare Caiman (Caiman yacare) fishing in flowing river, Pantanal, Brazil
Jacare Caiman (Caiman yacare) group on riverbank, Pantanal, Brazil
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Pete Oxford / Jacare Caiman (Caiman yacare) group on riverbank, Pantanal, Brazil
Jacare Caiman (Caiman yacare) thermoregulating on riverbank, Pantanal, Brazil
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Pete Oxford / Jacare Caiman (Caiman yacare) thermoregulating on riverbank, Pantanal, Brazil
Jacare Caiman (Caiman yacare) on riverbank, Pantanal, Brazil
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Pete Oxford / Jacare Caiman (Caiman yacare) on riverbank, Pantanal, Brazil
Jacare Caiman (Caiman yacare) on riverbank, Pantanal, Brazil
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Pete Oxford / Jacare Caiman (Caiman yacare) on riverbank, Pantanal, Brazil
Jacare Caiman (Caiman yacare) on riverbank, Pantanal, Brazil
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Pete Oxford / Jacare Caiman (Caiman yacare) on riverbank, Pantanal, Brazil
Jacare Caiman (Caiman yacare) thermoregulating on riverbank, Pantanal, Brazil
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Pete Oxford / Jacare Caiman (Caiman yacare) thermoregulating on riverbank, Pantanal, Brazil
Jacare Caiman (Caiman yacare) sunning on river bank, Pantanal, Brazil
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Luciano Candisani / Jacare Caiman (Caiman yacare) sunning on river bank, Pantanal, Brazil
Jacare Caiman (Caiman yacare) fishing in flowing river, Pantanal, Brazil
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Luciano Candisani / Jacare Caiman (Caiman yacare) fishing in flowing river, Pantanal, Brazil
Giant River Otter (Pteronura brasiliensis) an endangered species, eating fish in shallow water with Jacare Caiman (Caiman yacare) approaching, Pantanal, Brazil
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Theo Allofs / Giant River Otter (Pteronura brasiliensis) an endangered species, eating fish in shallow water with Jacare Caiman (Caiman yacare) approaching, Pantanal, Brazil
Aerial view of Parana river, before dam construction, Argentina / Paraguay border, South America
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Ross Couper-Johnston / Aerial view of Parana river, before dam construction, Argentina / Paraguay border, South America
Jacare Caiman (Caiman yacare) pair resting on river bank, Pantanal ecosystem, Brazil
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Claus Meyer / Jacare Caiman (Caiman yacare) pair resting on river bank, Pantanal ecosystem, Brazil
Paraguay River at sunset, Pantanal, Brazil
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Konrad Wothe / Paraguay River at sunset, Pantanal, Brazil
Common Water Hyacinth (Eichhornia crassipes) in Paraguay River, Pantanal, Brazil
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Konrad Wothe / Common Water Hyacinth (Eichhornia crassipes) in Paraguay River, Pantanal, Brazil
Cassauge River, Rio Paraguay, Pantanal, Mato Grosso, Brazil
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Konrad Wothe / Cassauge River, Rio Paraguay, Pantanal, Mato Grosso, Brazil
Jacare Caiman (Caiman yacare) group, fishing with open mouths in rushing river, at night, Pantanal, Brazil
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Tui De Roy / Jacare Caiman (Caiman yacare) group, fishing with open mouths in rushing river, at night, Pantanal, Brazil
Jacare Caiman (Caiman yacare) pair fishing the open mouths in rushing river, Pantanal, Brazil
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Tui De Roy / Jacare Caiman (Caiman yacare) pair fishing the open mouths in rushing river, Pantanal, Brazil
Jacare Caiman (Caiman yacare) group fishing with open mouths in rushing river, evening light, Pantanal, Brazil
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Tui De Roy / Jacare Caiman (Caiman yacare) group fishing with open mouths in rushing river, evening light, Pantanal, Brazil
Jacare Caiman (Caiman yacare) fishing in rushing river, Pantanal, Brazil
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Tui De Roy / Jacare Caiman (Caiman yacare) fishing in rushing river, Pantanal, Brazil
Jacare Caiman (Caiman yacare) trio fishing in flowing river, Pantanal, Brazil
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Tui De Roy / Jacare Caiman (Caiman yacare) trio fishing in flowing river, Pantanal, Brazil
Jacare Caiman (Caiman yacare), Cuiaba River, Pantanal, Brazil
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Daniel J. Cox / Jacare Caiman (Caiman yacare), Cuiaba River, Pantanal, Brazil
Jacare Caiman (Caiman yacare), Pantanal, Mato Grosso, Brazil
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Pete Oxford / Jacare Caiman (Caiman yacare), Pantanal, Mato Grosso, Brazil
Jacare Caiman (Caiman yacare) group on riverbank, Pantanal, Brazil
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Pete Oxford / Jacare Caiman (Caiman yacare) group on riverbank, Pantanal, Brazil
Jacare Caiman (Caiman yacare) group fishing with open mouths in rushing river at night, Pantanal, Brazil
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Tui De Roy / Jacare Caiman (Caiman yacare) group fishing with open mouths in rushing river at night, Pantanal, Brazil
Jacare Caiman (Caiman yacare) group fishing with open mouths in rushing river at night, Pantanal, Brazil
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Tui De Roy / Jacare Caiman (Caiman yacare) group fishing with open mouths in rushing river at night, Pantanal, Brazil
Jacare Caiman (Caiman yacare) group fishing with mouths open in rushing river, at night, Pantanal, Brazil
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Tui De Roy / Jacare Caiman (Caiman yacare) group fishing with mouths open in rushing river, at night, Pantanal, Brazil
Jacare Caiman (Caiman yacare) fishing with mouth open in rushing river, Pantanal, Brazil
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Tui De Roy / Jacare Caiman (Caiman yacare) fishing with mouth open in rushing river, Pantanal, Brazil
Jacare Caiman (Caiman yacare) fishing with mouth open, in rushing river, Pantanal, Brazil
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Tui De Roy / Jacare Caiman (Caiman yacare) fishing with mouth open, in rushing river, Pantanal, Brazil
Capybara (Hydrochoerus hydrochaeris) swimming, Little Paraguay River, Pocone, Brazil.
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Bruce Thomson / Capybara (Hydrochoerus hydrochaeris) swimming, Little Paraguay River, Pocone, Brazil.
Bare-faced curassow (Crax fasciolata) drinking at water's edge, Little Paraguay River, Pocone, Brazil.
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Bruce Thomson / Bare-faced curassow (Crax fasciolata) drinking at water's edge, Little Paraguay River, Pocone, Brazil.
Tapir (Tapirus terrestris) sniffing the air at dusk, Little Paraguay River, Pocone, Brazil.
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Bruce Thomson / Tapir (Tapirus terrestris) sniffing the air at dusk, Little Paraguay River, Pocone, Brazil.
RF - Giant Otters (Pteronura brasiliensis) in a lagoon off the Paraguay River, Taiama Reserve, western Pantanal, Brazil. (This image may be licensed either as rights managed or royalty free.)
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Nick Garbutt / RF - Giant Otters (Pteronura brasiliensis) in a lagoon off the Paraguay River, Taiama Reserve, western Pantanal, Brazil. (This image may be licensed either as rights managed or royalty free.)
Red-bellied piranha / red piranha (Pygocentrus nattereri / Serrasalmus nattereri) swimming, native to South America. Captive
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Philippe Clement / Red-bellied piranha / red piranha (Pygocentrus nattereri / Serrasalmus nattereri) swimming, native to South America. Captive