Paul Harcourt Davies / Emperor Moth (Saturnia pavoniella) male, on tree bark, Orvieto, Umbria, Italy, April

Paul Harcourt Davies,Paul Harcourt Davies / Lesser Horseshoe Bat (Rhinolophus hipposideros) hibernating in an Etruscan tomb, Podere Montecucco, near Orvieto, Umbria, Italy, November

Paul Harcourt Davies,Paul Harcourt Davies / Common Swallowtail butterfly (Papilio machaon) backlit showing wing structure. Podere Montecucco, Orvieto, Umbria, Italy, October

Paul Harcourt Davies / Short-horned grasshoppers (Calliptamus italicus) mating, showing sexual dimorphism, Pescia Romana, Tarquinia, Umbria, Italy, September

Paul Harcourt Davies / Newly emerged adult Two-tailed Pasha (Charaxes jasius) Podere Montecucco. Orvieto, Umbria, Italy August

Paul Harcourt Davies / Newly emerged adult Two-tailed Pasha butterfly (Charaxes jasius) Podere Montecucco. Orvieto, Umbria, Italy August

Paul Harcourt Davies,Paul Harcourt Davies / Emergence of Two-tailed Pasha butterfly (Charaxes jasius) Podere Montecucco. Orvieto, Umbria, Italy August. Sequence 6 of 9

Paul Harcourt Davies / Emergence of Two-tailed Pasha butterfly (Charaxes jasius) Podere Montecucco. Orvieto, Umbria, Italy August. Sequence 1 of 9

Paul Harcourt Davies,Paul Harcourt Davies / Martagon lily (Lilium martagonium) flower, Terminillo, Rieti, Lazio, Italy, July

Paul Harcourt Davies,Paul Harcourt Davies / Apollo butterfly (Parnasius apollo) feeding on flower, Mount Terminillo, Rieti, Lazio, Italy, July 2011

Paul Harcourt Davies / Alpine Snowbell (Soldanella alpina) in flower, above Madoona di Campiglio, Brenta Dolomites, Italy, June 2010

Paul Harcourt Davies / Floral colour in fields on the Piano Grande from Poppies (Papaver rhoeas), Cornflowers (Centaurea), Mustard (Brassica) and Mayweed (Anthemis), Umbria, Italy, June