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Sand bubble patterns created by Sand bubbler crabs (Scopimera inflata), Cape Hillsborough National Park, Queensland, Australia. September.
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Steven David Miller / Sand bubble patterns created by Sand bubbler crabs (Scopimera inflata), Cape Hillsborough National Park, Queensland, Australia. September.
Ural owl (Strix uralensis) female feeding her young with part of  a pellet it has regurgitated. Estonia, May.
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Sven Zacek / Ural owl (Strix uralensis) female feeding her young with part of a pellet it has regurgitated. Estonia, May.
Great horned owl (Bubo virginianus) chick coughing up a pellet, Southern California, USA, June.
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John Chan / Great horned owl (Bubo virginianus) chick coughing up a pellet, Southern California, USA, June.
Sand bubble patterns created by Sand bubbler crabs (Scopimera inflata), Cape Hillsborough National Park, Queensland, Australia. September.
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Steven David Miller / Sand bubble patterns created by Sand bubbler crabs (Scopimera inflata), Cape Hillsborough National Park, Queensland, Australia. September.
Brandt's Cormorant (Phalacrocorax penicillatus) pellet of worms coughed up by cormorant International Bird Rescue, Fairfield, California
01621940
Suzi Eszterhas / Brandt's Cormorant (Phalacrocorax penicillatus) pellet of worms coughed up by cormorant International Bird Rescue, Fairfield, California
Sparrowhawk (Accipter nisus) male regurgitating pellet, in forest, Pays de Loire, France
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Eric Medard / Sparrowhawk (Accipter nisus) male regurgitating pellet, in forest, Pays de Loire, France
Ural owl (Strix uralensis) female and chick at nest, female regurgitating pellet. Southern Estonia. May.
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Sven Zacek / Ural owl (Strix uralensis) female and chick at nest, female regurgitating pellet. Southern Estonia. May.
Close up of regurgitated pellet from great skua (Stercorarius skua) showing seabird prey remains like bones and feathers, Scotland, UK, May.
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Philippe Clement / Close up of regurgitated pellet from great skua (Stercorarius skua) showing seabird prey remains like bones and feathers, Scotland, UK, May.
Close up of regurgitated pellet from great skua (Stercorarius skua) showing seabird prey remains like bones and feathers, Scotland, UK, May.
01610558
Philippe Clement / Close up of regurgitated pellet from great skua (Stercorarius skua) showing seabird prey remains like bones and feathers, Scotland, UK, May.
European Bee-eater (Merops apiaster) regurgetating pellet, Andalucia, Spain
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Oscar Diez/ BIA / European Bee-eater (Merops apiaster) regurgetating pellet, Andalucia, Spain
Male Pied flycatcher (Ficedula hypoleuca) perched on a branch, disgorges pellet before flying out of frame, Carmarthenshire, Wales, UK, May.
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Dave Bevan / Male Pied flycatcher (Ficedula hypoleuca) perched on a branch, disgorges pellet before flying out of frame, Carmarthenshire, Wales, UK, May.
Barn owl (Tyto alba) pellets, contents of approximately six pellets containing skulls and bones of voles, mice and shrews, and cockchafer beetle. UK. March.
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Adrian Davies / Barn owl (Tyto alba) pellets, contents of approximately six pellets containing skulls and bones of voles, mice and shrews, and cockchafer beetle. UK. March.
Hawk Owl (Surnia ulula) regurgitating pellet before eating vole prey, Helsinki, Finland, December.
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Markus Varesvuo / Hawk Owl (Surnia ulula) regurgitating pellet before eating vole prey, Helsinki, Finland, December.
Great Horned Owl (Bubo virginianus) regurgitating pellet, Texas
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Alan Murphy/ BIA / Great Horned Owl (Bubo virginianus) regurgitating pellet, Texas
Short-eared owl (Asio flammeus) regurgitated pellet containing the remains of birds and mice, Scotland, UK, May
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Philippe Clement / Short-eared owl (Asio flammeus) regurgitated pellet containing the remains of birds and mice, Scotland, UK, May
Female Merlin (Falco columbarius) coughing up a pellet, Cairngorms National Park, Scotland, UK, June.
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SCOTLAND: The Big Picture / Female Merlin (Falco columbarius) coughing up a pellet, Cairngorms National Park, Scotland, UK, June.
Northern carmine bee-eater (Merops nubicus) regurgitating pellet, Selous Game Reserve, Tanzania.
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Charlie Summers / Northern carmine bee-eater (Merops nubicus) regurgitating pellet, Selous Game Reserve, Tanzania.
South polar skua (Stercorarius maccormicki) chick at nest site, regurgitates pellet, with parent in the foreground, Antarctica.
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Frederique Olivier / South polar skua (Stercorarius maccormicki) chick at nest site, regurgitates pellet, with parent in the foreground, Antarctica.
Rat bait, South Georgia Heritage Trust Rat Eradication Project, South Georgia 2011
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Ingo Arndt / Rat bait, South Georgia Heritage Trust Rat Eradication Project, South Georgia 2011
Wood sandpiper (Tringa glareola) regurgitating a pellet on the shore of a small lake in Southern Estonia.
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Sven Zacek / Wood sandpiper (Tringa glareola) regurgitating a pellet on the shore of a small lake in Southern Estonia.
Wood sandpiper (Tringa glareola) regurgitating a pellet on the shore of a small lake, Southern Estonia, May.
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Sven Zacek / Wood sandpiper (Tringa glareola) regurgitating a pellet on the shore of a small lake, Southern Estonia, May.
European roller (Coracias garrulus) regurgitating pellet perched on barbed wire fence, Alentero, Spain, April.
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Loic Poidevin / European roller (Coracias garrulus) regurgitating pellet perched on barbed wire fence, Alentero, Spain, April.
Common kingfisher (Alcedo atthis) regurgitating pellet, Le Teich, Gironde, France, September.
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Loic Poidevin / Common kingfisher (Alcedo atthis) regurgitating pellet, Le Teich, Gironde, France, September.
Reed warbler (Acrocephalus scirpaceus) feeding twelve day old Cuckoo (Cuculus canorus) chick at nest and removing faecal pellet, Norfolk, England, UK, May
01392959
David Tipling / 2020VISION / Reed warbler (Acrocephalus scirpaceus) feeding twelve day old Cuckoo (Cuculus canorus) chick at nest and removing faecal pellet, Norfolk, England, UK, May
European bee-eater (Merops apiaster) opening its beak wide before ejecting a pellet from it. Guerreiro, Castro Verde, Alentejo, Portugal, May..
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Roger Powell / European bee-eater (Merops apiaster) opening its beak wide before ejecting a pellet from it. Guerreiro, Castro Verde, Alentejo, Portugal, May..
Eagle owl (Bubo bubo) pellet containing remains of its prey, Luxembourg. March.
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Michel Poinsignon / Eagle owl (Bubo bubo) pellet containing remains of its prey, Luxembourg. March.
Common bee-eater (Merops apiaster) regurgitating pellet, Bouches du Rhone, France, May
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Loic Poidevin / Common bee-eater (Merops apiaster) regurgitating pellet, Bouches du Rhone, France, May
European Bee-eater (Merops apiaster) regurgitating pellet, France
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Walter Soestbergen / European Bee-eater (Merops apiaster) regurgitating pellet, France
Great tit {Parus major} flying from nest hole in tree trunk carrying faecal pellet, UK
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Laurent Geslin / Great tit {Parus major} flying from nest hole in tree trunk carrying faecal pellet, UK
Little owl (Athene noctua) coughing up a pellet, France, July
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Eric Medard / Little owl (Athene noctua) coughing up a pellet, France, July
Little owl (Athene noctua) coughing up a pellet, France, July
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Eric Medard / Little owl (Athene noctua) coughing up a pellet, France, July
Little owl (Athene noctua) during the day on a wall, coughing up a pellet, France, October
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Eric Medard / Little owl (Athene noctua) during the day on a wall, coughing up a pellet, France, October
Montagu's Harrier (Circus pygargus) male coughing up pellet, France, June
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Eric Medard / Montagu's Harrier (Circus pygargus) male coughing up pellet, France, June
Great tit (Parus major) flying flying from nest carrying faecal pellet, Lorraine, France, March.
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Michel Poinsignon / Great tit (Parus major) flying flying from nest carrying faecal pellet, Lorraine, France, March.
Snowy owl (Bubo scandiaca) male expelling pellet on the Canadian prairie, Saskatchewan, Canada, March, sequence 3/3
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Todd Mintz / Snowy owl (Bubo scandiaca) male expelling pellet on the Canadian prairie, Saskatchewan, Canada, March, sequence 3/3
Snowy owl (Bubo scandiaca) male expelling pellet on the Canadian prairie, Saskatchewan, Canada, March, sequence 2/3
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Todd Mintz / Snowy owl (Bubo scandiaca) male expelling pellet on the Canadian prairie, Saskatchewan, Canada, March, sequence 2/3
Snowy owl (Bubo scandiaca) male expelling pellet on the Canadian prairie, Saskatchewan, Canada, March, sequence 1/3
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Todd Mintz / Snowy owl (Bubo scandiaca) male expelling pellet on the Canadian prairie, Saskatchewan, Canada, March, sequence 1/3
Tawny owl (Strix aluco) regurgitated pellet, Berwickshire, Scotland, October
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Laurie Campbell / Tawny owl (Strix aluco) regurgitated pellet, Berwickshire, Scotland, October
Brown Rat (Rattus norvegicus) coming out of old pipe to feed on bait at whaling station for South Georgia Heritage Trust Rat Eradication Project, Grytviken, South Georgia Island
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Ingo Arndt / Brown Rat (Rattus norvegicus) coming out of old pipe to feed on bait at whaling station for South Georgia Heritage Trust Rat Eradication Project, Grytviken, South Georgia Island
Rat bait for South Georgia Heritage Trust Rat Eradication Project, South Georgia Island
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Ingo Arndt / Rat bait for South Georgia Heritage Trust Rat Eradication Project, South Georgia Island
Faecal pellet of Galapagos Giant Tortoise (Chelonoidis nigra) Wolf Volcano, Isabela Island, Galapagos
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Pete Oxford / Faecal pellet of Galapagos Giant Tortoise (Chelonoidis nigra) Wolf Volcano, Isabela Island, Galapagos
Short Eared Owl (Asio flammeus) coughing up an owl pellet. Lincolnshire, England, UK, February.
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Andy Sands / Short Eared Owl (Asio flammeus) coughing up an owl pellet. Lincolnshire, England, UK, February.
Short Eared Owl (Asio flammeus) coughing up an owl pellet. Lincolnshire, England, UK, February.
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Andy Sands / Short Eared Owl (Asio flammeus) coughing up an owl pellet. Lincolnshire, England, UK, February.
Ghost Crab (Ocypode sp) on beach surrounded by sand pellets which are by-products from the crab's feeding, Ecuador
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Murray Cooper / Ghost Crab (Ocypode sp) on beach surrounded by sand pellets which are by-products from the crab's feeding, Ecuador
Eastern Kingbird (Tyrannus tyrannus) ejecting a pellet of indigestible insect parts. Anahuac NWR, Texas, USA
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Gerrit Vyn / Eastern Kingbird (Tyrannus tyrannus) ejecting a pellet of indigestible insect parts. Anahuac NWR, Texas, USA
Regurgitated pellet of Tawny owl (Strix aluco) showing remains of prey, Sussex, England, UK
01275502
Adrian Davies / Regurgitated pellet of Tawny owl (Strix aluco) showing remains of prey, Sussex, England, UK
New Zealand / Sacred kingfisher (Todiramphus sancta vagans) ejecting pellet by regurgitating, Avon-Heathcote Estuary, Canterbury, New Zealand
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Tom Marshall / New Zealand / Sacred kingfisher (Todiramphus sancta vagans) ejecting pellet by regurgitating, Avon-Heathcote Estuary, Canterbury, New Zealand
Great Grey Shrike (Lanius excubitor) spitting up pellet, western Montana
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Donald M. Jones / Great Grey Shrike (Lanius excubitor) spitting up pellet, western Montana
Female Great Spotted Woodpecker (Dendrocopos major) flying from nest hole carrying faecal pellet, Lorraine, France
01193273
Michel Poinsignon / Female Great Spotted Woodpecker (Dendrocopos major) flying from nest hole carrying faecal pellet, Lorraine, France
Male Great Spotted Woodpecker (Dendrocopos major) flying from nest hole carrying faecal pellet, Lorraine, France
01193278
Michel Poinsignon / Male Great Spotted Woodpecker (Dendrocopos major) flying from nest hole carrying faecal pellet, Lorraine, France
Male Great Spotted Woodpecker (Dendrocopos major) flying from nest hole carrying faecal pellet, Lorraine, France
01193277
Michel Poinsignon / Male Great Spotted Woodpecker (Dendrocopos major) flying from nest hole carrying faecal pellet, Lorraine, France
Male Great Spotted Woodpecker (Dendrocopos major) flying from nest hole carrying faecal pellet, Lorraine, France
01193275
Michel Poinsignon / Male Great Spotted Woodpecker (Dendrocopos major) flying from nest hole carrying faecal pellet, Lorraine, France
Great Tit (Parus major) coming out of nest hole carrying faecal pellet, Lorraine, France
01193281
Michel Poinsignon / Great Tit (Parus major) coming out of nest hole carrying faecal pellet, Lorraine, France
Regurgitated bones of mammals from Harpy eagle nest (Harpia harpyja) Ecuador
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Pete Oxford / Regurgitated bones of mammals from Harpy eagle nest (Harpia harpyja) Ecuador
Fan-tailed warbler removes fecal pellet from nest {Cisticola juncidis} Spain
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Jose B. Ruiz / Fan-tailed warbler removes fecal pellet from nest {Cisticola juncidis} Spain
Orphean warbler removed fecal pellet from chick {Sylvia hortensis} Alicante, Spain
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Jose B. Ruiz / Orphean warbler removed fecal pellet from chick {Sylvia hortensis} Alicante, Spain
Common shrew bones {Sorex arabeus} found in pellet of Eurasian pygmy owl {Glaucidium passerinum}
01078905
Elio Della Ferrera / Common shrew bones {Sorex arabeus} found in pellet of Eurasian pygmy owl {Glaucidium passerinum}
Regurgitated Great horned owl pellets on taiga forest floor {Bubo virginianus} Denali NP, Alaska, USA.
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Philippe Clement / Regurgitated Great horned owl pellets on taiga forest floor {Bubo virginianus} Denali NP, Alaska, USA.
Owl pellet showing wing cases of beetles, Belgium
01140024
Philippe Clement / Owl pellet showing wing cases of beetles, Belgium
Tawny owl pellet {Strix aluco} showing remains of mouse skeleton, Belgium
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Philippe Clement / Tawny owl pellet {Strix aluco} showing remains of mouse skeleton, Belgium
Tawny owl pellet {Strix aluco} showing remains of mouse skeleton, Belgium
01140025
Philippe Clement / Tawny owl pellet {Strix aluco} showing remains of mouse skeleton, Belgium
Beetle {Byrrhus pilula} mimicing rabbit pellet. UK.
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Kim Taylor / Beetle {Byrrhus pilula} mimicing rabbit pellet. UK.
Kestrel {Falco tinnunculs} pellets containing mouse remains. UK.
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Kim Taylor / Kestrel {Falco tinnunculs} pellets containing mouse remains. UK.
Dor beetle {Typhaeus typhoeus} male with rabbit dropping / pellet. UK.
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Kim Taylor / Dor beetle {Typhaeus typhoeus} male with rabbit dropping / pellet. UK.
Harpy Eagle (Harpia harpyja) recently regurgitated pellet from fledged seven month old wild chick containing claws and fur from a Two-toed Sloth, Cuyabeno Reserve, Amazon rainforest, Ecuador
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Pete Oxford / Harpy Eagle (Harpia harpyja) recently regurgitated pellet from fledged seven month old wild chick containing claws and fur from a Two-toed Sloth, Cuyabeno Reserve, Amazon rainforest, Ecuador
Barn Owl (Tyto alba) owl pellets made of regurgitated bones and fur, Germany
00197433
Konrad Wothe / Barn Owl (Tyto alba) owl pellets made of regurgitated bones and fur, Germany
Regurgitated Pellets of Barn owl, Tawny owl, Short eared owl, Long eared owl and Little owl side-by-side for comparison
01101031
Mike Read / Regurgitated Pellets of Barn owl, Tawny owl, Short eared owl, Long eared owl and Little owl side-by-side for comparison
Common buzzard pellet {Buteo buteo} Beinn Eighe NNR, Scotland, UK
01096494
Niall Benvie / Common buzzard pellet {Buteo buteo} Beinn Eighe NNR, Scotland, UK
Contents of regurgitated Barn owl pellet {Tyto alba} UK. remains of woodmouse, vole, birds and short-tailed vole
01101024
Mike Read / Contents of regurgitated Barn owl pellet {Tyto alba} UK. remains of woodmouse, vole, birds and short-tailed vole
Lesser Horned Owl (Bubo magellanicus) pellet, Argentina
00595694
Agustin Esmoris / Lesser Horned Owl (Bubo magellanicus) pellet, Argentina
Eurasian Kestrel (Falco tinnunculus) juvenile regurgitating pellet at nest box, Hesse, Germany
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Duncan Usher / Eurasian Kestrel (Falco tinnunculus) juvenile regurgitating pellet at nest box, Hesse, Germany
Eurasian Kestrel (Falco tinnunculus) juvenile regurgitating pellet at nest box, Hesse, Germany
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Duncan Usher / Eurasian Kestrel (Falco tinnunculus) juvenile regurgitating pellet at nest box, Hesse, Germany
Northern Hawk Owl (Surnia ulula) pellets, Alaska
00584856
Michael Quinton / Northern Hawk Owl (Surnia ulula) pellets, Alaska
Mountain Pygmy-Owl (Glaucidium gnoma) regurgitating pellet, western Montana
00558077
Donald M. Jones / Mountain Pygmy-Owl (Glaucidium gnoma) regurgitating pellet, western Montana
Mountain Pygmy-Owl (Glaucidium gnoma) regurgitating pellet, western Montana
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Donald M. Jones / Mountain Pygmy-Owl (Glaucidium gnoma) regurgitating pellet, western Montana
Mountain Pygmy-Owl (Glaucidium gnoma) regurgitating pellet, western Montana, sequence 1 of 3
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Donald M. Jones / Mountain Pygmy-Owl (Glaucidium gnoma) regurgitating pellet, western Montana, sequence 1 of 3
American Pika (Ochotona princeps) collecting scat pellet from a Hoary Marmot (Marmota caligata), Glacier National Park, Montana
00537102
Sumio Harada / American Pika (Ochotona princeps) collecting scat pellet from a Hoary Marmot (Marmota caligata), Glacier National Park, Montana
Common Kingfisher (Alcedo atthis) pellet containing fish and dragonfly larva remains, Hessen, Germany
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Ingo Arndt / Common Kingfisher (Alcedo atthis) pellet containing fish and dragonfly larva remains, Hessen, Germany
Communal sphecid wasp (Trigonopsis sp) adding a pellet of mud to the outside of its communal nest on a tree in Amazonian rainforest, Brazil
01185143
Premaphotos / Communal sphecid wasp (Trigonopsis sp) adding a pellet of mud to the outside of its communal nest on a tree in Amazonian rainforest, Brazil
Barn owl (Tyto alba) regurgitated pellets showing close-up detail of mouse skull, hair and bones found inside. Belgium
01181526
Philippe Clement / Barn owl (Tyto alba) regurgitated pellets showing close-up detail of mouse skull, hair and bones found inside. Belgium
Barn owl (Tyto alba) regurgitated pellets showing close-up detail of mouse skull, hair and bones found inside. Belgium
01181527
Philippe Clement / Barn owl (Tyto alba) regurgitated pellets showing close-up detail of mouse skull, hair and bones found inside. Belgium
Barn owl  (Tyto alba) regurgitated pellets on hay bales in barn. Belgium
01181524
Philippe Clement / Barn owl (Tyto alba) regurgitated pellets on hay bales in barn. Belgium
Barn owl (Tyto alba) regurgitated pellets showing bones and skulls of mice. Belgium.
01181525
Philippe Clement / Barn owl (Tyto alba) regurgitated pellets showing bones and skulls of mice. Belgium.
Little owl (Athene noctua) coughing up pellet whilst perching on branch in meadow at dusk, Bedfordshire UK, June.
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Brian Bevan / Little owl (Athene noctua) coughing up pellet whilst perching on branch in meadow at dusk, Bedfordshire UK, June.
Little bee-eater (Merops pusillus) perched. The animal regurgitates a pellet of indigestible insect parts. Okavango Delta, Botswana.
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Neil Aldridge / Little bee-eater (Merops pusillus) perched. The animal regurgitates a pellet of indigestible insect parts. Okavango Delta, Botswana.
Little Bee-eater (Merops pusillus) regurgitating pellet, Samburu National Reserve, Kenya
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Steve Gettle / Little Bee-eater (Merops pusillus) regurgitating pellet, Samburu National Reserve, Kenya
Snowy owl (Bubo scandiaca) pellet, Canada January.
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Markus Varesvuo / Snowy owl (Bubo scandiaca) pellet, Canada January.
Bagworm moth caterpillar (Psychidae) defecating fecal pellet (frass) from its lower opening. The larvae lies hidden within its coccon casing, decorated with pieces of leaf to protect and disguise itse...
01704849
David Weiller / Bagworm moth caterpillar (Psychidae) defecating fecal pellet (frass) from its lower opening. The larvae lies hidden within its coccon casing, decorated with pieces of leaf to protect and disguise itse...
White-throated dipper (Cinclus cinclus) stretching wings then regurgitating food pellet, River Kennal, Cornwall, UK, January.
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Matt Roseveare / White-throated dipper (Cinclus cinclus) stretching wings then regurgitating food pellet, River Kennal, Cornwall, UK, January.
X-ray of a Pacific black duck (Anas superciliosa) showing ulna bone in wing shattered in 2 places from single pellet shot at Lake Leaghur in opening of hunting season. Bird rescued by Coalition Agains...
01689021
Doug Gimesy / X-ray of a Pacific black duck (Anas superciliosa) showing ulna bone in wing shattered in 2 places from single pellet shot at Lake Leaghur in opening of hunting season. Bird rescued by Coalition Agains...
Long-eared owl chick regurgitating pellet (Asio otus) perched in tree. Alsace. France.
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Eric Baccega / Long-eared owl chick regurgitating pellet (Asio otus) perched in tree. Alsace. France.
Owl pellet showing various rodent bones. Peak District National Park, Derbyshire, UK. June.
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Alex Hyde / Owl pellet showing various rodent bones. Peak District National Park, Derbyshire, UK. June.
Kingfisher (Alcedo atthis) female perched on branch regurgitating pellet, with bokeh and flare affect on river in the background, Lorraine, France, April
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Michel Poinsignon / Kingfisher (Alcedo atthis) female perched on branch regurgitating pellet, with bokeh and flare affect on river in the background, Lorraine, France, April
Filamentous fungus (Phycomyces blakesleeanus) beginning to form on mouse dropping. Hertfordshire, England, UK. November. Focus stacked image.
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Andy Sands / Filamentous fungus (Phycomyces blakesleeanus) beginning to form on mouse dropping. Hertfordshire, England, UK. November. Focus stacked image.
Bateleur (Terathopius ecaudatus) pair perched on branch, one coughing up pellet. Savuti, Chobe National Park, Botswana.
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Guy Edwardes / Bateleur (Terathopius ecaudatus) pair perched on branch, one coughing up pellet. Savuti, Chobe National Park, Botswana.
Researchers examining Raptor pellets to identify eaten prey. Part of a 60 year long-term study led by Fred Koning to monitor raptors and their nests in a 3,400 hectare area of coastal dunes. Near Amst...
01664054
Edwin Giesbers / Researchers examining Raptor pellets to identify eaten prey. Part of a 60 year long-term study led by Fred Koning to monitor raptors and their nests in a 3,400 hectare area of coastal dunes. Near Amst...