Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} groups walking along snow trail after heavy snow, Jigokudani, Nagano, Japan

Hanne & Jens Eriksen / Animal tracks in sand dune ripples, Liwa, UAE

Igor Shpilenok / Kamchatka brown bear (Ursus arctos beringianus) male sniffing the tracks of a female in snow, Kronotsky Zapovednik, Kamchatka, Far East Russia

Jouan & Rius / Antelope footprints in the white sand, Kalahari desert, South Africa

Anup Shah / Savanna grassland fire showing fire stopping at track which acts as fire break but has been crossed in places, Kenya, East Africa

Jack Dykinga / Boulder with zig-zag trails across a dry lake bed, sunset, Ubehebe Peak in distance, The Racetrack Playa, Death Valley National Park, California

Jack Dykinga / Saguaro Cacti (Carnegiea gigantea) silhouetted against night sky with star trails, time exposure for 5 hours after midnight, Cabeza Prieta National Wildlife Refuge, Arizona. New Years 2000

Jack Dykinga / Boulders with trails across a dry lake bed, sunrise, The Racetrack Playa, Death Valley National Park, California

Staffan Widstrand / Tracks of Polar bear in snow {Ursus maritimus} Baffin Island, Nunavut, North West Territories, Canada

Nick Garbutt / Rear view of male Bengal tiger walking {Panthera tigris tigris} Kanha NP, India - Winner in Wildlife Photographer of Year competition

Hermann Brehm / Oryx in the dunes of the Namib Desert, Namibia. (Oryx gazella)

Anup Shah / African lion (Panthera leo) walking along car tracks into distance, Serengeti National Park, Tanzania

Jean E. Roche / Weasel tracks showing hairs on feet {Mustela nivalis} Camargue, France

Yuri Shibnev / Footprints of Harvest mouse {Micromys minutus} in snow leading to print where bird landed and took mouse, far east Russia.

Yuri Shibnev / Tracks of European river otter in snow. {Lutra lutra} far east Russia

Igor Shpilenok / Kamchatka Brown bear (Ursus arctos beringianus) footprint in mud, Koryaksky Zapovednik, Russia.

Pete Cairns / Rock Ptarmigan {Lagopus mutus} tracks leading into Grampian Mountains, Scotland.

Anup Shah / Landscape with leopard on lookout rock{Panthera pardus}, Masai Mara Game Reserve, Kenya

Richard Du Toit / King cricket leaving tracks on sand {Mimnermidae} Namib Naukluft Park, Namibia

Richard Du Toit / Scorpion crossing sand dune leaving tracks {Scorpiones} Namib Naukluft P, Namibia

Pete Oxford / Tenebrionid beetle. Namibia (Onymacris bicolor). Desert.

DOC WHITE / Aerial view of two Beluga / White whale {Delphinapterus leucas} swimming at surface leaving ripples on calm sea surface, Canadian arctic

Geoff Simpson / Mountain hare track in snow {Lepus timidus} Peak District NP, UK

Sue Flood / Polar bear footprint in snow {Ursus maritimus} Churchill Canada

Ben Osborne / Imprint of Saltwater crocodile on sand {Crocodylus porosus} Queensland, Australia

Sue Daly / Lugworm cast {Arenicola marina} Jersey, Channel Isles, UK

Pete Oxford / Juvenile brown pelican {Pelecanus occidentalis} on beach, Espanola/Hood Is, Ecuador Book page 98

Mats Forsberg / Polar Bear tracks in snow, Svalbard , Norway

Hermann Brehm / Oryx (Oyrx gazella) in the dunes. Namib Desert, Namibia, Southern Africa

Neil Lucas / Horned viper {Cerastes cerastes} gliding sidewinding on sand dune, Morocco.

TJ Rich / Leopard {Panthera pardus} walking down sand dune Namibia, Southern Africa

Anup Shah / Tyre tracks across salt plain, Rann of Kutch desert, Gujarat, India

Richard Du Toit / Baboon spoor. Moremi reserve, Botswana, Southern Africa

David Tipling / Mountain hare (Lepus timidus) footprints in snow, with Spruce trees cloaked in snow Ruka Peak, Kuusamo, Finland. January.

David Tipling / Mountain hare (Lepus timidus) footprints in snow, with Spruce trees cloaked in snow Ruka Peak, Kuusamo, Finland. January.

David Tipling / Mountain hare (Lepus timidus) footprints in snow, among Spruce trees cloaked in snow Ruka Peak, Kuusamo, Finland. January.

David Tipling / Mountain hare (Lepus timidus) footprints in snow, with Spruce trees cloaked in snow Ruka Peak, Kuusamo, Finland. January.

Juan Carlos Munoz / Oak (Quercus sp) trees coming into leaf in field, tractor tracks in grassland, aerial view. La Gandara, Alto Ason, Soba Valley, Cantabria, Spain. May 2019.

Colin Varndell / Song thrush (Turdus philomelos) anvil with broken snail shells. Dorset, England, UK. April.

Steven David Miller / Rock art with stencilled hands, boomerangs and animal footprints . The Art Gallery, 62m of rock with 2000 stencils, engravings and paintings. Baloon Cave, Carnarvon Gorge, Carnarvon National Park, Que...

Juan Carlos Munoz / European river otter (Lutra lutra) footprints on riverbank mud. Ribeira de Piscos, Archaeological Park of the Coa Valley, Western Iberia, Portugal. April.

Yashpal Rathore / Bengal Tiger (Panthera tigris) foot print / pugmark on wet sand, Tadoba National Park, India

Gary Bell/ Oceanwide / Strawberry Land Hermit Crab (Coenobita perlatus) tracks on beach, Keeling Islands, Australia

Heather Angel / Feeding trails made by snail using radula strip with rows of tiny teeth on geen algal film on glass pane. Tropical butterfly house, Kunming, Yunnan Province, China.

Oriol Alamany / Red Fox (Vulpes vulpes) tracks in altiplano bear Gette, Spiti Valley, Cold Desert Biosphere Reserve, Himalaya, Himachal Pradesh, India, March

Yashpal Rathore / Kambala racing, a buffalo race which takes place on two parallel slushy tracks. Karnataka, India.

Neil Aldridge / Member of Botswana's Anti Poaching Unit keeps guard while a rhino monitor from the charity Rhino Conservation Botswana tracks a rhino in Okavango Delta, Botswana,. Rhino Conservation Botswana works ha...

Neil Aldridge / Director of Rhino Conservation Botswana Map Ives reads the footprints of a Black rhino (Diceros bicornis) in Okavango Delta, Botswana, where efforts have begun to rebuild the rhino populations the cou...

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) tracks in snow, Atka Bay, Queen Maud Land, Antarctica. October.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) tracks in snow, Atka Bay, Queen Maud Land, Antarctica. October.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) footprints in the snow. Atka Bay, Antarctica. July.

Stefan Christmann / Sea ice with hollows where Emperor penguin (Aptenodytes forsteri) breeding colony stood whilst huddling. Atka Bay, Antarctica. July.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) footprints in snow. Atka Bay, Antarctica. August.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) footprints on sea ice. Atka Bay, Antarctica. August.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) tracks in snow, during polar night. Atka Bay, Antarctica. June.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) foorprints in snow, during polar night. Atka Bay, Antarctica. June.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) foorprints in snow, during polar night. Atka Bay, Antarctica. June.

Sven Zacek / European elk (Alces alces), three standing together near frost covered Willow (Salix sp) bushes with Suur Emajogi River beyond. Alam-Pedja Nature Reserve, Tartumaa, Southern Estonia. January 2011.

Konrad Wothe / Firefly (Lamprohiza splendidula) light tracks in forest, Bavaria, Germany

Konrad Wothe / Firefly (Lamprohiza splendidula) light tracks in forest, Bavaria, Germany

Konrad Wothe / Firefly (Lamprohiza splendidula) light tracks in forest, Bavaria, Germany

Pal Hermansen / Ski tracks across a field, aerial view. Akershus, Norway. November 2017.

Axel Gomille / Wolf (Canis lupus) footprints, Saxony-Anhalt, Germany, August.

Axel Gomille / Wolf (Canis lupus) footprints in tire tracks, Saxony-Anhalt, Germany, August.

Axel Gomille / Wolf (Canis lupus) footprints, Saxony, Germany, July.

Axel Gomille / Wolf (Canis lupus) footprints, Saxony, Germany, August.

Ingo Arndt / Mountain Lion (Puma concolor) stepping into Guanaco (Lama guanicoe) tracks in snow during stalk, Torres del Paine National Park, Patagonia, Chile

Ingo Arndt / Mountain Lion (Puma concolor) tracks in snow, Torres del Paine National Park, Patagonia, Chile

Ingo Arndt / Mountain Lion (Puma concolor) tracks in snow, Torres del Paine National Park, Patagonia, Chile

Mark Carwardine / Blue whale (Balaenoptera musculus) flukeprint , aerial shot, Baja California, Mexico

Oriol Alamany / Footprint of Snow leopard (Panthera uncia) in snow, Spiti valley, Cold Desert Biosphere Reserve, Himalaya mountains, Himachal Pradesh, India, February

David Woodfall / Dinosaur footprints in Severn Estuary, Barry, Wales, UK, November.

David Woodfall / Long legged flies (Dolichopodidae) on puddle in foot print of Long Horn cattle. Knepp Wildland Project, formerly intensive farmland now turned to conservation and sustainable farming. Horsham, West Su...

Bruno D'Amicis / Marsican / Apennine brown bear (Ursus arctos marsicanus) claw marks on Beech (Fagus sylvatica) tree bark in old growth beech forest. Abruzzo, Lazio and Molise National Park / Parco Nazionale d'Abr...

Will Watson / Devonian sandstone with trace fossil. Tetrapod tracks created by a primitive four-limbed vertebrate circa 385 Million years ago. One of the oldest vertebrate tracks in the world. Valentia Island, Coun...

Alex Mustard / Tailing nudibranchs (Hypselodoris tryoni) on colourful sponges. This is a common behaviour in this species, where one individual tracks another, also known as queuing or tailgating. It may be relating...

Colin Varndell / Hazel nut (Corylus avellana) cases on forest floor, broken by grey squirrel. Dorset, UK August.

Luke Massey / Jaguar (Panthera onca) footprint, Brazil

Alain Dragesco-Joffe / Honey badger (Mellivora capensis) tracks in the sand, Tenere, Sahara, Niger.

Alain Dragesco-Joffe / Barbary sheep (Ammotragus lervia) footprints with ruler for scale. Sahara Desert, Nige.

Alain Dragesco-Joffe / Cheetah (Acionyx jubatus) tracks in the sand showing the typical characteristics of the cheetah - asymmetrical paws and non-retractible claws, Tenere, Sahara, Niger.

Alain Dragesco-Joffe / Striped hyena (Hyaena hyaena) footprints, showing typical larger rouder track of the front feet compared to back feet. Azaouak, Sahara Desert, Niger, .

Ashley Cooper / Children playing on tracks of railway bridge swept away by the January 2015 flooding leaving the tracks suspended in mid air, near Bangula. Malawi. March.

Ashley Cooper / Children playing on tracks of railway bridge swept away by the January 2015 flooding leaving the tracks suspended in mid air, near Bangula. Malawi. March.

Ashley Cooper / Derwent Water completely frozen over with cycle tracks, Keswick, Lake District, England, UK, December 2010.

Ashley Cooper / Derwent Water completely frozen over with cycle tracks, Keswick, Lake District, England, UK, December 2010.

Ashley Cooper / Tracks left by the bark beetles that have killed the tree. As climate change causes warmer winter temperatures, bark beetles are migrating further north killing millions of acres of forest. Alberta,...

Ashley Cooper / Bridge to cross river which has dried up during the drought which lasted from 1996-2011. Victoria, Australia, February 2010.

Ashley Cooper / Truck hauling freshly cut timber for biofuel in Grizedale forest, Lake District, England, UK, May 2013.

Ashley Cooper / A forwarder, a specialist machine for cutting timber for biofuel in Grizedale Forest, Lake District, England, UK

Ashley Cooper / Trams on the street in Hong Kong, China. February 2010.

Ashley Cooper / Trams on the street in Hong Kong, China. February 2010.

Neil Aldridge / Member of the Botswana Defence Force keeping guard while a rhino monitor from Rhino Conservation Botswana tracks an adult rhino in northern Botswana. September 2017. Highly Commended in the Natural Wo...

Sue Flood / Emperor penguins (Aptenodytes forsteri) tracks from tobogganing, Antarctica.

Ann & Steve Toon / Brown hyaena (Hyaena brunnea) footprints in sand, Namib desert, Namibia.

Martin Willis / Purple-crowned Fairywren (Malurus coronatus) male calling to female, Lawnhill River, Queensland, Australia

Martin Willis / Purple-crowned Fairywren (Malurus coronatus) male calling to female, Lawnhill River, Queensland, Australia

Eric Sohn Joo Tan / Purple-crowned Fairywren (Malurus coronatus) male, Western Australia, Australia

Mike Potts / Purple Crowned Fairy Wren (Malurus coronatus) ringed as part of a study, adult male, Mornington NP Kimberly Region, Western Australia, July

Mike Potts / Purple Crowned Fairy Wren (Malurus coronatus) ringed as part of a study, adult male, Mornington NP Kimberly Region, Western Australia, July

Mike Potts / Purple Crowned Fairy Wren (Malurus coronatus) ringed as part of a study, adult male, Mornington NP Kimberly Region, Western Australia, July

Mike Potts / Purple Crowned Fairy Wren (Malurus coronatus) Mornington NP, Kimberly Region, Western Australia, July

Martin Willis / Purple-crowned Fairywren (Malurus coronatus) male, Lawnhill River, Queensland, Australia

Etienne Littlefair / Purple-crowned fairywren (Malurus coronatus) perched on branch, Victoria River, Northern Territory, Australia.

Martin Willis / Purple-crowned Fairywren (Malurus coronatus) female, Northern Territory, Australia

Martin Willis / Purple-crowned Fairywren (Malurus coronatus) male, Northern Territory, Australia