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Bioluminescence from glowing plankton in sea tide line on beach, with stars above and ship lights on horizon, Vaadhoo Island, Raa Atoll, Maldives, Indian Ocean, October 2010
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Doug Perrine / Bioluminescence from glowing plankton in sea tide line on beach, with stars above and ship lights on horizon, Vaadhoo Island, Raa Atoll, Maldives, Indian Ocean, October 2010
Reef manta ray (Manta alfredi) shoal filter feeding on plankton concentrated by monsoon currents in Raa Atoll, Vandhoo Thila, Maldives. October.
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Tui De Roy / Reef manta ray (Manta alfredi) shoal filter feeding on plankton concentrated by monsoon currents in Raa Atoll, Vandhoo Thila, Maldives. October.
Bioluminescence from glowing plankton in sea tide line on beach, with stars above and ship lights on horizon, Vaadhoo Island, Raa Atoll, Maldives, Indian Ocean, October 2010
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Doug Perrine / Bioluminescence from glowing plankton in sea tide line on beach, with stars above and ship lights on horizon, Vaadhoo Island, Raa Atoll, Maldives, Indian Ocean, October 2010
Reef manta ray (Manta alfredi) filter feeding on plankton. Madhivafaru Reef, Raa Atoll, Maldives
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Tui De Roy / Reef manta ray (Manta alfredi) filter feeding on plankton. Madhivafaru Reef, Raa Atoll, Maldives
Strawberry hermit crab (Coenobita perlatus) on white coral sand beach, Small unhinhabited coral islet in Raa Atoll lagoon, Maldives
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Tui De Roy / Strawberry hermit crab (Coenobita perlatus) on white coral sand beach, Small unhinhabited coral islet in Raa Atoll lagoon, Maldives
Strawberry land hermit crab (Coenobita perlatus).  On white coral sand beach, Small unhinhabited coral islet in Raa Atoll lagoon, Maldives
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Tui De Roy / Strawberry land hermit crab (Coenobita perlatus). On white coral sand beach, Small unhinhabited coral islet in Raa Atoll lagoon, Maldives
House crow (Corvus splendens) small unhinhabited coral islet in atoll lagoon, Raa Atol, Maldives.
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Tui De Roy / House crow (Corvus splendens) small unhinhabited coral islet in atoll lagoon, Raa Atol, Maldives.
Maldives tropical reef community. Mixed fish species schooling over mostly dead or dying coral reef following several bleaching events, the latest in 2016. Kottefaru Beyru, Raa Atoll, Maldives.
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Tui De Roy / Maldives tropical reef community. Mixed fish species schooling over mostly dead or dying coral reef following several bleaching events, the latest in 2016. Kottefaru Beyru, Raa Atoll, Maldives.
Remora (Remora remora), freeswimming near Manta Ray, Dhikkurendho Reef, Raa Atoll, Maldives.
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Tui De Roy / Remora (Remora remora), freeswimming near Manta Ray, Dhikkurendho Reef, Raa Atoll, Maldives.
Pennant coralfish (Heniochus acuminatus), over mostly dead or dying coral reef following several bleaching events, Vadhoo Thila, Raa Atoll, Maldives.
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Tui De Roy / Pennant coralfish (Heniochus acuminatus), over mostly dead or dying coral reef following several bleaching events, Vadhoo Thila, Raa Atoll, Maldives.
Reef manta ray (Manta alfredi) Visiting cleaning station to invite small reef fish to clean it of skin parasites. Neyo Beyru, Raa Atoll, Maldives
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Tui De Roy / Reef manta ray (Manta alfredi) Visiting cleaning station to invite small reef fish to clean it of skin parasites. Neyo Beyru, Raa Atoll, Maldives
Reef manta ray (Manta alfredi) tourists swimming with mantas filter feeding, Dhikkurendho Reef, Raa Atoll, Maldives
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Tui De Roy / Reef manta ray (Manta alfredi) tourists swimming with mantas filter feeding, Dhikkurendho Reef, Raa Atoll, Maldives
Reef manta rays (Manta alfredi) filter feeding on plankton, Dhikkurendho Reef, Raa Atoll, Maldives
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Tui De Roy / Reef manta rays (Manta alfredi) filter feeding on plankton, Dhikkurendho Reef, Raa Atoll, Maldives
Reef manta rays (Manta alfredi) filter feeding on plankton, Dhikkurendho Reef, Raa Atoll, Maldives
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Tui De Roy / Reef manta rays (Manta alfredi) filter feeding on plankton, Dhikkurendho Reef, Raa Atoll, Maldives
Reef manta rays (Manta alfredi) filter feeding on plankton, Dhikkurendho Reef, Raa Atoll, Maldives
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Tui De Roy / Reef manta rays (Manta alfredi) filter feeding on plankton, Dhikkurendho Reef, Raa Atoll, Maldives
Reef manta rays (Manta alfredi) tourists swimming with mantas filter feeding on plankton, Dhikkurendho Reef, Raa Atoll, Maldives
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Tui De Roy / Reef manta rays (Manta alfredi) tourists swimming with mantas filter feeding on plankton, Dhikkurendho Reef, Raa Atoll, Maldives
Reef manta rays (Manta alfredi) filter feeding on plankton, Dhikkurendho Reef, Raa Atoll, Maldives
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Tui De Roy / Reef manta rays (Manta alfredi) filter feeding on plankton, Dhikkurendho Reef, Raa Atoll, Maldives
Reef manta rays (Manta alfredi) filter feeding on plankton, Dhikkurendho Reef, Raa Atoll, Maldives
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Tui De Roy / Reef manta rays (Manta alfredi) filter feeding on plankton, Dhikkurendho Reef, Raa Atoll, Maldives
Reef manta rays (Manta alfredi) filter feeding on plankton, Dhikkurendho Reef, Raa Atoll, Maldives
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Tui De Roy / Reef manta rays (Manta alfredi) filter feeding on plankton, Dhikkurendho Reef, Raa Atoll, Maldives
Reef manta rays (Manta alfredi) filter feeding on plankton, Dhikkurendho Reef, Raa Atoll, Maldives
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Tui De Roy / Reef manta rays (Manta alfredi) filter feeding on plankton, Dhikkurendho Reef, Raa Atoll, Maldives
Reef manta ray (Manta alfredi)  filter feeding on plankton.  Madhivafaru Reef, Raa Atoll, Maldives
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Tui De Roy / Reef manta ray (Manta alfredi) filter feeding on plankton. Madhivafaru Reef, Raa Atoll, Maldives
Reef manta rays (Manta alfredi) filter feeding on plankton  Hanifaru Bay, Raa Atoll, Maldives
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Tui De Roy / Reef manta rays (Manta alfredi) filter feeding on plankton Hanifaru Bay, Raa Atoll, Maldives
Reef manta rays (Manta alfredi) filter feeding in atoll passes and lagoons Hanifaru Bay, Raa Atoll, Maldives
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Tui De Roy / Reef manta rays (Manta alfredi) filter feeding in atoll passes and lagoons Hanifaru Bay, Raa Atoll, Maldives
Reef manta rays (Manta alfredi) filter feeding in atoll passes and lagoons Hanifaru Bay, Raa Atoll, Maldives
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Tui De Roy / Reef manta rays (Manta alfredi) filter feeding in atoll passes and lagoons Hanifaru Bay, Raa Atoll, Maldives
Reef manta rays (Manta alfredi) filter feeding in atoll passes and lagoons Hanifaru Bay, Raa Atoll, Maldives
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Tui De Roy / Reef manta rays (Manta alfredi) filter feeding in atoll passes and lagoons Hanifaru Bay, Raa Atoll, Maldives
Reef manta rays (Manta alfredi) filter feeding in atoll passes and lagoons Hanifaru Bay, Raa Atoll, Maldives
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Tui De Roy / Reef manta rays (Manta alfredi) filter feeding in atoll passes and lagoons Hanifaru Bay, Raa Atoll, Maldives
Reef manta ray (Manta alfredi) barrel-roll feeding, Hanifaru Bay, Raa Atoll, Maldives
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Tui De Roy / Reef manta ray (Manta alfredi) barrel-roll feeding, Hanifaru Bay, Raa Atoll, Maldives
Reef manta rays (Manta alfredi) barrel-roll feeding, Hanifaru Bay, Raa Atoll, Maldives
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Tui De Roy / Reef manta rays (Manta alfredi) barrel-roll feeding, Hanifaru Bay, Raa Atoll, Maldives
Reef manta ray (Manta alfredi) filter feeding on plankton. Vandhoo Thila, Raa Atoll, Maldives
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Tui De Roy / Reef manta ray (Manta alfredi) filter feeding on plankton. Vandhoo Thila, Raa Atoll, Maldives
Reef manta rays (Manta alfredi) filter feeding on plankton. Vandhoo Thila, Raa Atoll, Maldives
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Tui De Roy / Reef manta rays (Manta alfredi) filter feeding on plankton. Vandhoo Thila, Raa Atoll, Maldives
Reef manta rays (Manta alfredi) filter feeding on plankton. Vandhoo Thila, Raa Atoll, Maldives
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Tui De Roy / Reef manta rays (Manta alfredi) filter feeding on plankton. Vandhoo Thila, Raa Atoll, Maldives
Reef manta ray (Manta alfredi) filter feeding on plankton. Vandhoo Thila, Raa Atoll, Maldives
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Tui De Roy / Reef manta ray (Manta alfredi) filter feeding on plankton. Vandhoo Thila, Raa Atoll, Maldives
Reef manta rays (Manta alfredi) filter feeding on plankton. Vandhoo Thila, Raa Atoll, Maldives
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Tui De Roy / Reef manta rays (Manta alfredi) filter feeding on plankton. Vandhoo Thila, Raa Atoll, Maldives