Tony Wu / Twinspot snapper (Lutjanus bohar) shoal rushing up to spawn, releasing a cloud of milky white sperm and eggs in the water column. Caroline Islands, Palau, November.

Tony Wu / Twinspot snapper (Lutjanus bohar) engaged in a mass spawning aggregation early in the morning. Caroline Islands, Palau. April. Winner of Underwater category, Wildlife Photographer of the Year 2016.

Doug Perrine / Barjacks (Caranx ruber), Yellowtail snapper (Ocyurus chrysurus), and Sergeant major damselfish (Abudefduf saxatilis) swimming around wreck of bow gun on C-58 shipwreck off Puerto Morelos, near Cancun,...

Alex Mustard / Portrait of pair of Bohar snappers (Lutjanus bohar) in their spawning aggregation. Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea.

Alex Mustard / Portrait of Bohar snappers (Lutjanus bohar) in its spawning aggregation. Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea.

Alex Mustard / Glasseye snapper (Heteropriacanthus cruentatus) head portrait with a cleaning Goby (Gobiosoma genie) East End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea.

Georgette Douwma / Shoal of Bohar / Twinspot snapper {Lutjanus bohar} Ras Mohammed, Red Sea, Egypt

Georgette Douwma / Kaleidoscopic image of Bigeye snappers (Lutjanus lutjanus) with soft corals. West Papua, Indonesia.

Sergio Hanquet / Glasseye snapper (Heteropriacanthus cruentatus), headshot. Tenerife, Canary Islands.

Sergio Hanquet / Glasseye snapper (Heteropriacanthus cruentatus) portrait. Tenerife, Canary Islands.

Shane Gross / Schoolmaster snapper (Lutjanus apodus) shoal sheltering amongst Red mangrove (Rhizophora mangle) roots. Caribbean Sea off Gardens of the Queen National Park, Cuba.

Alex Mustard / Coral grouper (Cephalopholis miniata) waits in ambush, hiding against a seafan (Annella sp.) hunting silversides (Atherinidae), with schooling bigeye snapper (Lutjanus lutjanus) behind. Misool, Raja A...

Shane Gross / Schoolmaster snapper (Lutjanus apodus) in red mangrove(Rhizophora mangle) and turtlegrass (thalassia testudinum) habitats, Eleuthera Island, Bahamas.

David Fleetham / Blacktail snapper (Lutjanus fulvus) in coral reef. Kauai, Hawaii.

Georgette Douwma / Bigeye snapper (Lutjanus lutjanus) school over coral reef. Triton Bay, West Papua, Indonesia.

Linda Pitkin / Bigeye snappers (Lutjanus lutjanus) schooling in coral reef gully, Triton Bay, near Kaimana, West Papua, Indonesia

Linda Pitkin / Slender pinjalo (Pinjalo lewisi) fish schooling, Triton Bay, near Kaimana, West Papua, Indonesia

Linda Pitkin / Slender pinjalo (Pinjalo lewisi) fish schooling, Triton Bay, near Kaimana, West Papua, Indonesia

Linda Pitkin / Slender pinjalo (Pinjalo lewisi) fish schooling, Triton Bay, near Kaimana, West Papua, Indonesia

Linda Pitkin / Slender Pinjalo (Pinjalo lewisi) fish schooling, Triton Bay, near Kaimana, West Papua, Indonesia, February 2017

David Fleetham / Adult Two-spot Red Snapper (Lutjanus bohar) among other reef fish. Bequ Lagoon, Fiji.

Alex Mustard / School of Bohar snapper (Lutjanus bohar) swimming in formation along a coral reef. These fish are usually solitary and aggregate each summer in the Red Sea to spawn. Ras Mohammed National Park, Sinai...

Alex Mustard / Blackspot snapper (Lutjanus ehrenbergii). Beacon Rock, Sinai, Egypt. Red Sea.

Alex Mustard / School of Bohar snapper (Lutjanus bohar) in open water above a coral reef. These fish are usually solitary and aggregate each summer in the Red Sea to spawn. Shark Reef, Ras Mohammed, Sinai, Egypt. Re...

Alex Mustard / School of Bohar snapper (Lutjanus bohar) in open water close to a coral reef. These fish are usually solitary and aggregate each summer in the Red Sea to spawn. Shark Reef, Ras Mohammed, Sinai, Egypt....

Alex Mustard / School of Bohar snapper (Lutjanus bohar) in open water close to a coral reef. Shark Reef, Ras Mohammed, Sinai, Egypt. Red Sea.

Alex Mustard / School of Bohar snapper (Lutjanus bohar) close to a coral reef. These fish are usually solitary and aggregate each summer in the Red Sea to spawn. Shark Reef, Ras Mohammed, Sinai, Egypt. Red Sea.

Constantinos Petrinos / Emperor red snaper (Lutjanus sebae) sub-adult. Lembeh Strait, North Sulawesi, Indonesia.

Georgette Douwma / Bigeye snapper (Lutjanus lutjanus) shoal in coral reef scenery including Soft corals (Dendronephthya sp), West Papua, Indonesia.

Georgette Douwma / Bigeye snappers (Lutjanus lutjanus) shoal swimming in coral reef including Black coral (Antipathes dichotoma), West Papua, Indonesia.

Georgette Douwma / Bigeye snapper (Lutjanus lutjanus) shoal in coral reef, West Papua, Indonesia.

Alex Mustard / Bohar snappers (Lutjanus bohar) breaking up into smaller groups to spawn close to full moon. The fish swim short arches above the main group as they release clouds of gametes (eggs from the female, sp...

Alex Mustard / Large school of Bohar snappers (Lutjanus bohar) swimming in a tight, fast moving circular school over the reef, just before swimming into open water to spawn. Shark City, Ulong, Rock Islands, Palau. T...

Alex Mustard / Bohar snappers (Lutjanus bohar) breaking up into smaller groups to spawn close to full moon. The fish swim short arches above the main group as they release clouds of gametes (eggs from the female, sp...

Alex Mustard / Bohar snappers (Lutjanus bohar) breaking up into smaller groups to spawn close to full moon. The fish swim short arches above the main group as they release clouds of gametes (eggs from the female, sp...

Claudio Contreras / Glasseye Snapper (Heteropriacanthus cruentatus), San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve (Socorro Islands), Pacific Ocean, Western Mexico, March

Linda Pitkin / Bluestripe snappers (Lutjanus kasmira) and Bigeye Snappers (Lutjanus lutjanus) in mixed-species school. Mioskon, Dampier Strait, Raja Ampat, West Papua, Indonesia.

Linda Pitkin / Bigeye snappers (Lutjanus lutjanus) in mixed-species school with Bluestripe snappers (Lutjanus kasmira) Mioskon, Dampier Strait, Raja Ampat, West Papua, Indonesia.

Alex Mustard / Bohar snapper (Lutjanus bohar) school at Ras Mohammed, Shark Reef, Ras Mohammed, Sinai, Egypt. Red Sea.

Tony Wu / Humpback red snapper (Lutjanus gibbus) juvenile, Kokopo, Papua New Guinea.

Pascal Kobeh / Red snappers / Two-spot red snappers (Lutjanus bohar) mass shoal of fish spawning, Palau, Philippine Sea. A few black and white snappers (Macolor niger) have crawled into the school to eat the eggs.

Alex Mustard / School of Bohar snappers (Lutjanus bohar) above a coral reef. These fish are usually solitary but have gathered in the summer in the Red Sea for spawning. Shark Reef, Ras Mohammed Marine Park, Sinai,...

Alex Mustard / Bohar snappers (Lutjanus bohar) breaking up into smaller groups to spawn. The fish swim short arches above the main group as they release clouds of gametes. Shark City, Ulong, Rock Islands, Palau. T...

Georgette Douwma / Bigeye snappers (Lutjanus lutjanus) shoal, Andaman Sea, Thailand.

Georgette Douwma / Bigeye snappers (Lutjanus lutjanus) shoal, Mabul, Malaysia.

Alex Mustard / Streaming school of Bigeye snappers (Lutjanus lutjanus) swims across a reef filled with fish. Andiamo, Daram Islands, Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea, Pacific Ocean.

Alex Mustard / Bohar snapper (Lutjanus bohar) school with diver, Ras Mohammed, Ras Mohammed, Sinai, Egypt. Red Sea.

Pascal Kobeh / Mexican goatfish (Mulloidichthys dentatus) and Blue-and-gold snappers (Lutjanus viridis) in a feeding frenzy over Triggerfish eggs, Baja California peninsula, Mexico. Sea of Cortez.

Pete Oxford / Northern Red Snapper (Lutjanus campechanus) and Barracuda (Sphyraena sp) being sold in market, Suva, Viti Levu, Fiji

Alex Mustard / Large school of Bohar snappers (Lutjanus bohar) which have gathered in the summer in the Red Sea for spawning. Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea.

Alex Mustard / School of Bohar snappers (Lutjanus bohar) Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea.

Alex Mustard / Portrait of the, normally solitary Bohar snapper (Lutjanus bohar) in an annual spawning aggregation in June. Ras Mohammed Marine Park, Sinai, Egypt. Red Sea.

Alex Mustard / The vertical reef wall at Shark Reef, Ras Mohammed, with Scalefin anthias (Pseudanthias squamipinnis) and soft corals (Dendronephthya spp.) Yellowsaddle goatfish (Parupeneus cyclostomus) and school of...

Alex Mustard / School of Bohar snappers (Lutjanus bohar), moving along the face of reef. These fish are up to 80cm in length and normally solitary, but have gathered in summer in spawning aggregation. Shark Reef, Ra...

Alex Mustard / Large school of Bohar snappers (Lutjanus bohar) which have gathered in the summer in the Red Sea for spawning. Each fish is between 60 and 80cm long. Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt...

Alex Mustard / School of large Bohar snappers (Lutjanus bohar) gathered in spawning aggregation off Ras Mohammed, Sinai, Egypt. Red Sea.

Alex Mustard / Large school of Bohar snappers (Lutjanus bohar) gather with smaller shoal of Batfish (Platax orbicularis) in open water off Ras Mohammed at the tip of Sinai, Egypt. These schools gather only in summer...

Alex Mustard / School of Batfish (Platax orbicularis) in front of Bohar snappers (Lutjanus bohar) as the wall at Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea.

Michael Pitts / Bluelined snapper (Lutjanus coeruleolineatus) Red Sea.

Franco Banfi / Shoal of Humpback red snapper (Lutjanus gibbus) Maldives, Indian Ocean

Shibai Xiao / Mangrove red snapper (Lutjanus Argentimaculatus) captrive underwater with two fisherman on bank looking down into water, Hainan Island, South China Sea, China. No release available.

Pete Oxford / Caught Red Snapper (Lutjanus campechanus). Mahahual Dock, Mahahual, South Yucatan Peninsula, Mexico.

Juergen Freund / Schooling Pinjalo snapper (Pinjalo lewisi) West New Britain, Papua New Guinea.

Juergen Freund / Juvenile red emperor snapper (Lutjanus sebae), swimming among sea urchins. Lembeh Strait, North Sulawesi, Indonesia.

Georgette Douwma / Mangrove jack / red snapper (Lutjanus argentimaculatus) swimming along coral wall with soft corals and crinoids, Rinca, Indonesia

GEORGETTE DOUWMA / Brown-striped red snapper (Lutjanus vitta) with soft corals. Andaman Sea, Thailand.

Tim Laman / Snapper fish {Lutjanidae} and rich invertebrate life growing underwater on Red mangrove roots {Rhizophora mangle} in the Belize Cays, Tunicate Cove, Belize.

Tim Laman / Giant anemone {Condylactis gigantea}, Snapper fish {Lutjanidae}, sponges, tunicates and other invertebrates amongst the roots of Red Mangrove trees {Rhizophora mangle} in the Belize Cays, Tunicate Cov...

Tim Laman / Snapper fish {Lutjanidae}, sponges, tunicates and other invertebrates amongst the roots of Red Mangrove trees {Rhizophora mangle} in the Belize Cays, Tunicate Cove, Belize.

Tim Laman / Snapper fish {Lutjanidae}, sponges, tunicates and other invertebrates amongst the roots of Red Mangrove trees {Rhizophora mangle} in the Belize Cays, Tunicate Cove, Belize.

GEORGETTE DOUWMA / Red emperor fish / snapper (Lutjanus sebae) feeding on a Grey bonnet (Phalium glaucum). Lembeh Strait, North Sulawesi, Indonesia.

GEORGETTE DOUWMA / Twinspot / Bohar snapper (Lutjanus bohar), Red Sea, Egypt

GEORGETTE DOUWMA / Diver with shoal of Bohar / Twinspot snapper {Lutjanus bohar} Ras Mohammed, Red Sea, Egypt

GEORGETTE DOUWMA / Shoal of Bohar / Twinspot snapper {Lutjanus bohar} Ras Mohammed, Red Sea, Egypt

GEORGETTE DOUWMA / Emperor snapper {Lutjanus sebae} juvenile sheltering in a False fire urchin {Astropyga radiata} Lembeh Strait, North Sulawesi, Indonesia.

GEORGETTE DOUWMA / Shoal of One spot snapperfish {Lutjanus lutjanus} Egypt, Red Sea

Gary Bell/ Oceanwide / Yellow-banded Snapper (Lutjanus adetii) and Emperor Red Snapper (Lutjanus sebae), Heron Island, Great Barrier Reef, Queensland, Australia

David Fleetham / Freediver with a speared Mangrove jack / red snapper (Lutjanus argentimaculatus) Palau, Micronesia. Model released.

Noemie Stroh / Red snapper (Lutjanus bohar) aggregation during mating season, Ras Mohamed National Park, Red Sea, Egypt. July.

Noemie Stroh / One-spot snapper (Lutjanus monostigma) schooling above coral reef, Sataya, Red Sea, Egypt. August.

Alex Mustard / Schools of fish, including Bohar snappers (Lutjanus bohar), Yellow-spotted trevally (Carangoides orthogrammus) and Bigeye trevally (Caranx sexfasciatus) gathering adjacent to a colourful coral reef wi...

Alex Mustard / Vermilion rockfish (Sebastes miniatus) juvenile, portrait, Vancouver Island, British Columbia, Canada, Queen Charlotte Strait, Pacific Ocean.

Henley Spiers / Red snapper (Lutjanus bohar) in foreground and other reef fish in background including Napoleon wrasse (Cheilinus undulatus) feeding on discarded fish, Misool, Raja Ampat, Indonesia, Pacific Ocean

Sergio Hanquet / African striped grunt (Parapristipoma octolineatum) shoal swimming under rock with a Glasseye Snapper (Heteropriacanthus cruentatus) amongst them , Tenerife, Canary Islands, Atlantic Ocean.

Georgette Douwma / Yellow tropical reef fish composite image on black background, Falcula butterflyfish (Chaetodon falcula), Yellow boxfish (Ostracion cubicus), Golden butterflyfish (Chaetodon semilarvatus), Red Sea clo...

Georgette Douwma / Yellow tropical reef fish composite image on black background, Sailfin tang (Zebrasoma veliferum) juvenile, Pajama cardinalfish (Sphaeramia nematoptera), Three-spot angelfish (Apolemichthys trimaculat...

Henley Spiers / Bigeye snapper (Lutjanus lutjanus) school swimming in formation, Misool Marine Reserve, Raja Ampat, West Papua, Pacific Ocean.

Noemie Stroh / Blackspot snappers (Lutjanus ehrenbergii) schooling on a coral reef, Sataya, Red Sea, Egypt, August.

David Fleetham / Slate pencil sea urchin (Heterocentrotus mammillatus) on a reef with Black triggerfish (Melichthys niger) and a Bluestripe snapper (Lutjanus kasmira) swimming above, Maui, Hawaii, Pacific Ocean.

Tony Wu / Reef manta ray (Mobula alfredi) swimming with a Twinspot snapper (Lutjanus bohar), Okinawa, Japan, Pacific Ocean.

David Fleetham / Glasseye snapper (Heteropriacanthus cruentatus) in a cavern during the day, Hawaii.

David Fleetham / Blacktail snapper (Lutjanus fulvus) with school of shoulderbar Soldierfish (Myripristis kuntee) Hawaii.

Shane Gross / Grey snapper fish (Lutjanus griseus) hunting Silversides (Atherinomorus lacunosus) among red mangrove (Rhizophora mangle) roots, Eleuthera, Bahamas.

Sirachai Arunrugstichai / Large colonies of Gorgonian sea fan (Melithea sp.) with an unusual white colouration, not related to coral bleaching. With a school of Bigeye snapper (Lutjanus lutjanus) at Hin Khao, Andaman coast, Sa...

Sirachai Arunrugstichai / Healthy Staghorn corals (Acropora sp.) with various reef fishes, including Moon wrasses (Thalassoma lunare), and Threespot dascyllus (Dascyllus trimaculatus) and a large school of Bigeye snappers (Lut...