Nick Garbutt / Silhouette of Red-fronted brown lemur (Eulemur rufus) leaping across gap in trees by moonlight, cathemeral behaviour, Ranomafana National Park, south east Madagascar, November

Alex Hyde / Red fronted brown lemurs {Lemur fulvus rufus} drinking from pool, Kirindy forest, West Madagascar.

Alex Hyde / Close up of fore foot of Red fronted brown lemur {Lemur fulvus rufus}, Kirindy forest, West Madagascar

Edwin Giesbers / Red fronted brown lemur (Lemur fulvus rufus) Madagascar

Pete Oxford / Red fronted lemur. Female. Madagascar, Kirindy forest.

Suzi Eszterhas / Red-fronted brown lemur (Eulemur rufifrons) infant clinging to mother, Berenty Private Reserve, Madagascar.

Suzi Eszterhas / Red-fronted brown lemur (Eulemur rufifrons) Berenty Private Reserve, Madagascar

Suzi Eszterhas / Red-fronted brown lemur (Eulemur rufifrons) Berenty Private Reserve, Madagascar

Chien Lee / Red-fronted Brown Lemur (Eulemur fulvus rufus), Andasibe, Madagascar

Chien Lee / Red-fronted Brown Lemur (Eulemur fulvus rufus), Andasibe, Madagascar

Chien Lee / Red-fronted Brown Lemur (Eulemur fulvus rufus), Andasibe, Madagascar

Chien Lee / Red-fronted Brown Lemur (Eulemur fulvus rufus), Andasibe, Madagascar

Bernard Castelein / Red-fronted lemur (Eulemur rufifrons) male, Kirindy Forest, Madagascar.

Bernard Castelein / Red-fronted lemur (Eulemur rufifrons) male in tree, Kirindy Forest, Madagascar.

Bernard Castelein / Red-fronted lemur (Eulemur rufifrons) male in tree, Kirindy Forest, Madagascar.

Bernard Castelein / Red-fronted lemur (Eulemur rufifrons) male, Kirindy Forest, Madagascar.

Bernard Castelein / Red-fronted lemur (Eulemur rufifrons) males on forest floor,, Kirindy Forest, Madagascar.

Bernard Castelein / Red-fronted lemur (Eulemur rufifrons) male in tree, Kirindy Forest, Madagascar.

Enrique Lopez-Tapia / Common brown lemur (Eulemur fulvus) and Red-fronted brown lemurs (Eulemur rufus) in tree, Andasibe-Mantadia National Park, Madagascar.

Konrad Wothe / Rufous Brown Lemur (Eulemur rufus) male climbing tree, Andasibe Mantadia National Park, Madagascar, Africa

Konrad Wothe / Red-fronted Brown Lemur (Eulemur fulvus rufus) female with young, Berenty Reserve, Madagascar

Konrad Wothe / Red-fronted Brown Lemur (Eulemur fulvus rufus) male in tree, Isalo National Park, Madagascar

Konrad Wothe / Red-fronted Brown Lemur (Eulemur fulvus rufus) female in tree, Andasibe Mantadia National Park, Madagascar

Konrad Wothe / Red-fronted Brown Lemur (Eulemur fulvus rufus) male in tree, Andasibe Mantadia National Park, Madagascar

Konrad Wothe / Red-fronted Brown Lemur (Eulemur fulvus rufus) male on brach, Andasibe Mantadia National Park, Madagascar

Mark Carwardine / Red fronted brown lemur {Eulemur fulvus} Berenty Private Reserve, southern Madagascar

Loic Poidevin / Red frontal brown lemur (Eulemur fulvus rufus) drinking from a forest stream, Madagascar.

Loic Poidevin / Red frontal brown / Southern red-fronted brown lemur (Eulemur fulvus rufus) sitting on branch in forest canopy, Madagascar.

Loic Poidevin / Red frontal brown / Southern red-fronted brown lemur (Eulemur fulvus rufus) two cuddled up together sitting on branch in forest canopy, Madagascar.

Alex Hyde / Red fronted brown lemur {Lemur fulvus rufus} with baby, looking down from tree, Kirindy forest, West Madagascar

Alex Hyde / Red fronted brown lemur {Lemur fulvus rufus}, Kirindy forest, West Madagascar

Alex Hyde / Red fronted brown lemurs {Lemur fulvus rufus} drinking from pool, Kirindy forest, West Madagascar.

Alex Hyde / Red fronted brown lemur {Lemur fulvus rufus}, portrait, Kirindy forest, West Madagascar

Alex Hyde / Red fronted brown lemur {Lemur fulvus rufus} grooming foot, Kirindy forest, West Madagascar.

Alex Hyde / Red fronted brown lemur {Lemur fulvus rufus} looking down from tree, Kirindy forest, West Madagascar.

Nick Garbutt / Female Red fronted brown lemur (Lemur fulvus rufus) grooming, Kirindy forest, West Madagascar

Pete Oxford / Female Red fronted brown lemur (Lemur fulvus rufus) licking fingers, Kirindy, West Madagascar

Pete Oxford / Female Red fronted brown lemur (Lemur fulvus rufus) Kirindy forest, West Madagascar

Pete Oxford / Red fronted brown lemurs (Lemur fulvus rufus) walking along track in Western dry forest, West Madagascar

Pete Oxford / Male Red fronted brown lemur (Lemur fulvus rufus) Kirindy forest, West Madagascar

Pete Oxford / Female Red fronted brown lemur (Lemur fulvus rufus) licking baby, Kirindy Reserve, West Madagascar

Pete Oxford / Male Red fronted lemur (Lemur fulvus rufus) feeding on tamarind, Berenty Private Reserve, Madagascar

Pete Oxford / Red fronted lemur (Lemur fulvus rufus) female sitting on branch, Berenty private reserve, Madagascar

Vincent Grafhorst / Red-fronted Brown Lemur (Eulemur fulvus rufus) with identification collar, Kirindy Forest, Madagascar

Fiona Rogers / Red-fronted brown lemur (Eulemur rufus) female feeding on a chameleon. Berenty Private Reserve, Madagascar. Oct 2008.

Fiona Rogers / Red-fronted brown lemur (Eulemur rufus) female feeding on a chameleon. Berenty Private Reserve, Madagascar. Oct 2008.

Anup Shah / Ring-tailed Lemur female (Lemur catta) in confrontation with Red-fronted Brown Lemur female (Eulemur rufus) over food. Berenty Private Reserve, Madagascar. Oct 2008.

Anup Shah / Ring-tailed Lemur (Lemur catta) female in confrontation over food with female Red-fronted Brown Lemur (Eulemur rufus) Berenty Private Reserve, Madagascar. Oct 2008.

Anup Shah / Red-fronted Brown Lemur (Eulemur rufus) male and female sitting. Berenty Private Reserve, Madagascar. Oct 2008.

Anup Shah / Red-fronted Brown Lemur (Eulemur rufus) male and female sitting. Berenty Private Reserve, Madagascar. Oct 2008.

Anup Shah / Red-fronted Brown Lemur (Eulemur rufus) male portrait. Berenty Private Reserve, Madagascar. Oct 2008.

Anup Shah / Red-fronted Brown Lemur (Eulemur rufus) female feeding on cactus. Berenty Private Reserve, Madagascar. Oct 2008.

Anup Shah / Red-fronted Brown Lemur (Eulemur rufus) female walking. Berenty Private Reserve, Madagascar. Oct 2008.

Anup Shah / Red-fronted Brown Lemur (Eulemur rufus) male sitting. Berenty Private Reserve, Madagascar. Oct 2008.

Anup Shah / Red-fronted brown lemur (Eulemur rufus) female standing alert. Berenty Private Reserve, Madagascar. Oct 2008.

Edwin Giesbers / Red fronted brown lemur (Lemur fulvus rufus) sitting on branch, Madagascar

Edwin Giesbers / Red fronted brown lemur (Lemur fulvus rufus) silhouette, Berenty, Western Madagascar

Edwin Giesbers / Red fronted brown lemur (Lemur fulvus rufus) portrait, Madagascar

Edwin Giesbers / Red fronted brown lemur (Lemur fulvus rufus) sitting in tree, Madagascar

Edwin Giesbers / Red fronted brown lemur (Lemur fulvus rufus) walking along leaf covered branch, Madagascar

Edwin Giesbers / Red fronted brown lemur (Lemur fulvus rufus) baby peering out from behind mother, Madagascar

Edwin Giesbers / Red fronted brown lemur (Lemur fulvus rufus) baby peering out from behind mother, Madagascar

Edwin Giesbers / Red fronted brown lemur (Lemur fulvus rufus) portrait, Madagascar

Edwin Giesbers / Red fronted brown lemur (Lemur fulvus rufus) portrait, Madagascar

Edwin Giesbers / Red fronted brown lemur (Lemur fulvus rufus) portrait, Madagascar

Edwin Giesbers / Red fronted brown lemur (Lemur fulvus rufus) sitting on branch, Madagascar

Edwin Giesbers / Red fronted brown lemur (Lemur fulvus rufus) baby peering out from behind mother with finger in its mouth, Madagascar

Edwin Giesbers / Red fronted brown lemur (Lemur fulvus rufus) sitting on branch, Madagascar

Edwin Giesbers / Red fronted brown lemur (Lemur fulvus rufus) portrait, Madagascar

Edwin Giesbers / Red fronted brown lemur (Lemur fulvus rufus) portrait, Madagascar

Edwin Giesbers / Red fronted brown lemur (Lemur fulvus rufus) sitting in tree, Madagascar

Inaki Relanzon / Red-fronted brown lemur (Eulemur rufus / Lemur fulvus rufus) Kirindy forest, Madagascar

Inaki Relanzon / Red fronted brown lemur (Eulemur rufus / Lemur fulvus rufus) feeding on fruit in tree, Berenty Private Reserve, South Madagascar

Jouan & Rius / Red fronted brown lemur (Eulemur fulvus rufus) eating papaya fruit, Madagascar

Konrad Wothe / Red-fronted Brown Lemur (Eulemur fulvus rufus) male, Ranomafana National Park, Madagascar

Konrad Wothe / Red-fronted Brown Lemur (Eulemur fulvus rufus) male, Madagascar

Konrad Wothe / Red-fronted Brown Lemur (Eulemur fulvus rufus) male, Ranomafana National Park, Madagascar

Nick Garbutt / Red fronted lemur (Eulemus / Lemur fulvus rufus} infant nestling on mother's thigh, Ranomafana National Park, Madagascar

Pete Oxford / Red-fronted Brown Lemur (Eulemur fulvus rufus) male portrait, Berenty Reserve, southern Madagascar

Nick Garbutt / Baby Red fronted lemur {Lemur fulvus rufus} portrait.

Pete Oxford / Red fronted lemur pair with young (Lemur fulvus rufus) Kirindy forest, Madagascar

Pete Oxford / Red fronted lemur {Lemur fulvus rufus} western dry forest, Madagascar

Pete Oxford / Red fronted lemur {Lemur fulvus rufus} female with collar for research, Kirindy forest, Western Madagascar,

Pete Oxford / Red fronted lemur {Lemur fulvus rufus} female eating fruit, Berenty, Western Madagascar,

Pete Oxford / Red fronted lemur {Lemur fulvus rufus} male, Berenty, Western Madagascar,

Pete Oxford / Red fronted lemur {Lemur fulvus rufus} female in tree, Kirindy forest, Western Madagascar,

Pete Oxford / Male red-fronted lemur {Lemur fulvus rufus}. Berenty primate reserve, Madagascar.

Pete Oxford / Red fronted brown lemurs {Lemur fulvus rufus} inKirindy forest, W Madagascar

Pete Oxford / Female Red fronted brown lemur {Lemur fulvus rufus} Kirindy forest, W Madagascar

Lynn M Stone / Red Fronted Lemurs (Lemur fulvus rufus) sitting on branch, captive, from Madagascar.

Lynn M Stone / Red Fronted Lemur carrying young (Lemur fulvus rufus) captive, from Madagascar.

Lynn M Stone / Red Fronted Lemur with young (Lemur fulvus rufus) captive, from Madagascar.

Pete Oxford / Red fronted lemur {Lemur fulvus rufus} male, Kirindy Forest, Madagascar

Pete Oxford / Red fronted brown lemur, Berenty, Madagascar

Pete Oxford / Red-fronted brown lemur (lemur fulvus rufus} with young, Berenty, Madagascar

Pete Oxford / Female Red fronted brown lemur with baby, Kirindy Forest, Madagascar