Heike Odermatt / Brunnich's Guillemot (Uria lomvia) nesting colony on cliff, Alkefjellet, Hinlopen Strait, Svalbard, Norway

Danny Green / Brunnich's guillemot (Uria lomvia) colony in mist, many birds in flight and nesting on cliffs. Svalbard, Norway, July 2018.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony huddled together under the Southern Lights, Aurora australis, Atka Bay, Antarctica. September.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony, males huddling whilst incubating eggs. Atka Bay, Antarctica. July. Winner of the Portfolio category of the Wildlife Photographer of the Year Awards 2019.

Marie Read / Arctic terns (Sterna paradisaea), two in flight over nesting colony in field of buttercups (Ranunculus sp.), Keflavik, Iceland. Digital canvas expansion.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony with chicks, Atka Bay, Queen Maud Land, Antarctica. October.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony with with creche of huddling chicks, Atka Bay, Queen Maud Land, Antarctica. October.

Alex Hyde / Colonies of King penguins (Aptenodytes patagonicus) and Southern Elephant Seals (Mirounga leonina) on beach at St. Andrews Bay, South Georgia. November.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) breeding colony, males huddling on sea ice during polar night, incubating eggs, ice shelf in background. Atka Bay, Antarctica. July 2017.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony huddled during polar night, males incubating eggs. Atka Bay, Antarctica. July 2017.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony, males huddling whilst incubating eggs. Atka Bay, Antarctica. July.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony, males huddling whilst incubating eggs. Atka Bay, Antarctica. July.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) males in breeding colony huddling during winter storm. Atka Bay, Antarctica. July.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony huddling, males incubating eggs, on sea ice under full moon in polar night. Atka Bay, Antarctica. July 2017.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) breeding colony on sea ice, ice shelf in background. Atka Bay, Antarctica. August.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) males in breeding colony incubating eggs, huddling during winter storm. Penguins on outside repositioning themselves to gain protection from weather. Atka Bay, A...

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) breeding colony, mainly males brooding eggs, huddled together on sea ice. Atka Bay, Antarctica. June.

Stefan Christmann / Cameraman Lindsay McCrae filming Emperor penguin (Aptenodytes forsteri) colony for BBC Dynasties Penguin programme. Chicks are on sea ice whilst parents fish at sea. Atka Bay, Antarctica. December 201...

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) group sliding over ice, returning to form breeding colony. Atka Bay, Antarctica. April.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) group sliding over sea ice, returning to form breeding colony. Iceberg in background. Atka Bay, Antarctica. April 2017.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) walking across crack in sea ice during return to breeding colony. Atka Bay, Antarctica. April.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) group sliding across sea ice to form breeding colony. Atka Bay, Antarctica. April 2017.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) sliding across sea ice in journey to breeding colony. Sastrugi in ice formed by wind erosion. Atka Bay, Antarctica. April.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) forming breeding colony at start of winter. Atka Bay, Antarctica. April.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri) breeding colony, adults brooding chicks. Atka Bay, Antarctica. September 2017.

Stefan Christmann / Emperor penguin (Aptenodytes forsteri), breeding colony with individuals in foreground walking to / from open water. Atka Bay, Antarctica. September 2017.

Philippe Clement / Common guillemots (Uria aalge) and Razorbills (Alca torda) swimming and fishing at sea, Fowlsheugh RSPB Reserve, Aberdeenshire, Scotland, UK, May.

Philippe Clement / Razorbill (Alca torda) looking around on sea cliff top at seabird colony in spring, Fowlsheugh RSPB Reserve, Aberdeenshire, Scotland, UK, May

Tui De Roy / Waved albatross (Phoebastria irrorata) group of three on nest, Punta Suarez, Espanola Island, Galapagos

Frederique Olivier / Close-up of a group of Emperor penguin (Aptenodytes forsteri) chicks huddled together in a blizzard, covered in snow, Adelie Land, Antarctica, January.

Frederique Olivier / Emperor penguin (Aptenodytes forsteri) chicks huddled together during a blizzard, Adelie Land, Antarctica, January.

Frederique Olivier / Group of Adelie penguins (Pygoscelis adeliae) destroying an egg with a chick inside, Adelie Land, Antarctica, January.

Frederique Olivier / Close-up of a Black browed albatross (Thalassarche melanophris) preening its chick, chick looks up panting, thermoregulating, West Point Island, Falkland Islands.

Frederique Olivier / Southern rockhopper penguin (Eudyptes chrysocome) at nest in breeding colony, stretching and knocking chick over, New Island, Falkland Islands.

Frederique Olivier / Southern rockhopper penguin (Eudyptes chrysocome) preening chick then itself, New Island, Falkland Islands.

Frederique Olivier / Black browed albatross (Thalassarche melanophris) chick in nest, interacting with a Southern rockhopper penguin (Eudyptes chrysocome) chick, New Island, Falkland Islands.

Frederique Olivier / Chinstrap penguin (Pygoscelis antarcticus) moving a feather in its nest, Aitcho Island, South Shetland Islands.

Frederique Olivier / Chinstrap penguin (Pygoscelis antarcticus) adding a stick to its nest and fighting with a neighbouring pair, Aitcho Island, South Shetland Islands.

Frederique Olivier / Panning shot of a Chinstrap penguin (Pygoscelis antarcticus) breeding colony, Aitcho Island, South Shetland Islands.

Peter Bassett / Pair of Gentoo penguins (Pygoscelis papua) courting and attempting to mate, Neko Harbour, Andvord Bay, Graham Land, Antarctica.

Peter Bassett / Panning shot of a King penguin (Aptenodytes patagonicus) breeding colony, Salisbury Plain, South Georgia.

Peter Bassett / King penguin (Aptenodytes patagonicus), Salisbury Plain, South Georgia.

Peter Bassett / King penguins (Aptenodytes patagonicus) preening in breeding colony, with mountains in the background, Salisbury Plain, South Georgia.

Peter Bassett / King penguins (Aptenodytes patagonicus) entering and leaving the sea near their breeding colony, Salisbury Plain, South Georgia.

Peter Bassett / King penguins (Aptenodytes patagonicus) in a breeding colony, with mountains in the background, Salisbury Plain, South Georgia.

Peter Bassett / King penguins (Aptenodytes patagonicus) walking towards and entering the sea near their breeding colony, Salisbury Plain, South Georgia.

Frederique Olivier / Pair of King penguins (Aptenodytes patagonicus) regurgitating and calling with chick nearby, Macquarie Island, Sub-Antarctic Australia.

Tui De Roy / Snares Crested penguin (Eudyptes robustus) group walking up granite slope into Oleander forest, Station Point colony. Snares Island, New Zealand Sub-Antarctic Islands.

Frederique Olivier / Adelie penguin (Pygoscelis adeliae) colony, with icebergs and pack ice in the background, Antarctica.

Frederique Olivier / Adelie penguin (Pygoscelis adeliae) displaying in the middle of a colony, Antarctica.

Ingo Arndt / Seabird colony nesting on cliff face, Lange Anna, Helgoland, Germany

Claudio Contreras / Great Frigatebird (Fregata minor) departing from nest, San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Isl...

Neil Aldridge / African penguins (Spheniscus demersus) preening themselves on rocks at the Boulders Beach colony near Cape Town, South Africa.

Hiroya Minakuchi / King Penguin (Aptenodytes patagonicus) colony, South Georgia Island

Mark MacEwen / Snares island crested penguin (Eudyptes robustus) colony in forest, Snares Island, New Zealand.

Mark MacEwen / Snare's island crested penguin (Eudyptes robustus) colony on the coast, high angle view, Snares Island, New Zealand.

David Tipling / Northern gannet (Sula bassana) colony, Hermaness, Unst, Shetland Scotland, UK, April.

Espen Bergersen / Common guillemot (Uria aalge) flock, circling outside the breeding colony at Hornoya birdcliff, Finnmark, Norway.

Chris & Monique Fallows / King penguin (Aptenodytes patagonicus) colony at sunrise. Grytviken, South Georgia Island.

Chris & Monique Fallows / King penguin (Aptenodytes patagonicus) colony with storm approaching. Grytviken, South Georgia Island.

Chris & Monique Fallows / King penguin (Aptenodytes patagonicus) and Southern elephant seal (Mirounga leonina) colony at sunrise. Grytviken, South Georgia Island.

Stefan Christmann / Emperor Penguin (Aptenodytes forsteri) colony huddling for warmth, Queen Maud Land, Antarctica

Stefan Christmann / Emperor Penguin (Aptenodytes forsteri) colony huddling for warmth, Queen Maud Land, Antarctica

Mark Jones / Snares-crested penguin (Eudyptes robustus) group climbing steep route to colony. Snares Island, New Zealand Subantarctic Islands. Endemic. Vulnerable species.

Bryan and Cherry Alexander / Emperor penguins (Aptenodytes forsteri) colony, with Snow petrel (Pagodroma nivea) flying past and dark sky. Atka Bay. Antarctica

Cheryl-Samantha Owen / Cape gannet (Morus capensis) colony, Bird Island Nature Reserve, Lambert's Bay, West Coast, South Africa.

Andrew Parkinson / Gannet (Morus bassanus) adult, backlit by the morning sun, coming in to land in colony, Shetland Islands, Scotland, UK. September

Andrew Parkinson / Cliff top view of a gannet (Morus bassanus) colony. Shetland Islands, Scotland, UK, July.

Barrie Britton / King Penguin (Aptenodytes patagonicus) colony, with mountains reflected in the ocean, St Andrew's Bay, South Georgia. Photograph taken on location for the BBC Frozen Planet series, October 2009.

Erlend Haarberg / Northern gannet (Morus bassanus) colony on a sea stack and seabird cliff, Langanes peninsula, Iceland, July

Peter Cairns / 2020VISION / Silhouette of Razorbill (Alca torda) against sunset, flapping wings. June 2010. Photographer quote: 'A cacophonous seabird colony on a summers evening is just spell-binding. Throw in a decent sunset a...

Pete Oxford / Black-browed Albatross (Thalassarche melanophrys) nesting colony, Steeple Jason Island, Falkland Islands

Pete Oxford / Black-browed Albatross (Thalassarche melanophrys) nesting colony, Steeple Jason Island, Falkland Islands

Orsolya Haarberg / Atlantic puffin (Fratercula arctica). Langanes peninsula, Iceland. May 2008

Orsolya Haarberg / Razorbill (Alca torda), mating. Langanes peninsula, Iceland. May 2008

Orsolya Haarberg / Common guillemot (Uria aalge) and razorbills (Alca torda). Langanes peninsula, Iceland. May 2008

Orsolya Haarberg / Northern gannet (Morus bassanus) colony, courtship, Seabird cliff, Langanes peninsula, Iceland, July 2008

Orsolya Haarberg / Atlantic puffin (Fratercula arctica). Ltrabjarg seabird cliff, Iceland. July 2008

Orsolya Haarberg / Atlantic puffin (Fratercula arctica). Ltrabjarg seabird cliff, Iceland. July 2008

Orsolya Haarberg / Atlantic puffin (Fratercula arctica). Ltrabjarg seabird cliff, Iceland. July 2008

Orsolya Haarberg / Atlantic puffin (Fratercula arctica). Ltrabjarg seabird cliff, Iceland. July 2008

Orsolya Haarberg / Northern gannet (Morus bassanus) colony, Seabird cliff, Langanes peninsula, Iceland, July 2008

Orsolya Haarberg / Northern gannet (Morus bassanus) colony, Seabird cliff, Langanes peninsula, Iceland, July 2008

Orsolya Haarberg / Northern gannet (Morus bassanus) colony, Seabird cliff, Langanes peninsula, Iceland, July 2008

Yva Momatiuk & John Eastcott / Gentoo Penguin (Pygoscelis papua) parent huddling with two chicks, Booth Island, Antarctica

Thomas Mangelsen / King Penguin (Aptenodytes patagonicus) group of downy chicks, Fortuna Bay, South Georgia

Flip de Nooyer / Gentoo Penguin (Pygoscelis papua) two downy chicks standing together in nesting colony, Antarctica

Flip de Nooyer / Black-browed Albatross (Thalassarche melanophrys) chicks on tower nests, South Georgia Island

Tui De Roy / Salvin's Albatross (Thalassarche salvini) returning en masse to crowded nesting colony at sunset, Proclamation Island, Bounty Islands, New Zealand

Tui De Roy / Salvin's Albatross (Thalassarche salvini) returning to crowded nesting colony at sunset, Proclamation Island, Bounty Islands, New Zealand

Yva Momatiuk & John Eastcott / Atlantic Puffin (Fratercula arctica) adults congregate in colony on coastal cliff during summer breeding season, Newfoundland, Canada

Pete Oxford / Emperor Penguin (Aptenodytes forsteri) pair, Flutter EP Rookery, Cape Darnley, Australian Antarctic Territory, Antarctica

Gordon Court / Adelie Penguin (Pygoscelis adeliae) gives a warning call when under threat, Cape Bird, Ross Island, Antarctica

Tui De Roy / Campbell Albatross (Thalassarche impavida) portrait of two month old chick awaiting parent's feeding, Bull Rock, North Cape Colony, Campbell Island, New Zealand

Tui De Roy / Campbell Albatross (Thalassarche impavida) coming in to land, Bull Rock Cape Colony, Campbell Island, New Zealand

Tui De Roy / King Penguin (Aptenodytes patagonicus) group swimming underwater offshore from breeding colony, Macquarie Island, sub-Antarctica Australia