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African elephant (Loxodonta africana) on plains under stormy sky, Marakele National Park, Waterberg Biosphere, South Africa
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Andy Rouse / African elephant (Loxodonta africana) on plains under stormy sky, Marakele National Park, Waterberg Biosphere, South Africa
Burchell's Zebra (Equus burchellii) pair nuzzling, Rietvlei Nature Reserve, Gauteng, South Africa
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Richard Du Toit / Burchell's Zebra (Equus burchellii) pair nuzzling, Rietvlei Nature Reserve, Gauteng, South Africa
Burchell's Zebra (Equus burchellii) yawning, Rietvlei Nature Reserve, Gauteng, South Africa
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Richard Du Toit / Burchell's Zebra (Equus burchellii) yawning, Rietvlei Nature Reserve, Gauteng, South Africa
Cattle Egret (Bubulcus ibis) on Burchell's Zebra (Equus burchellii), Rietvlei Nature Reserve, Gauteng, South Africa
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Richard Du Toit / Cattle Egret (Bubulcus ibis) on Burchell's Zebra (Equus burchellii), Rietvlei Nature Reserve, Gauteng, South Africa
Burchell's Zebra (Equus burchellii) males fighting, Rietvlei Nature Reserve, Gauteng, South Africa
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Richard Du Toit / Burchell's Zebra (Equus burchellii) males fighting, Rietvlei Nature Reserve, Gauteng, South Africa
Burchell's Zebra (Equus burchellii) pair, Rietvlei Nature Reserve, Gauteng, South Africa
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Richard Du Toit / Burchell's Zebra (Equus burchellii) pair, Rietvlei Nature Reserve, Gauteng, South Africa
Burchell's Zebra (Equus burchellii) males fighting, Rietvlei Nature Reserve, Gauteng, South Africa
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Richard Du Toit / Burchell's Zebra (Equus burchellii) males fighting, Rietvlei Nature Reserve, Gauteng, South Africa
Mozambique Spitting Cobra (Naja mossambica) ejecting venom, Kruger, South South Africa, Controlled conditions
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Guy Edwardes / Mozambique Spitting Cobra (Naja mossambica) ejecting venom, Kruger, South South Africa, Controlled conditions
Mozambique Spitting Cobra (Naja mossambica) ejecting venom Kruger, South Africa, Controlled conditions.
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Guy Edwardes / Mozambique Spitting Cobra (Naja mossambica) ejecting venom Kruger, South Africa, Controlled conditions.
African elephant (Loxodonta africana) male tusker walking down road photographed from vehicle, iMfolozi National Park, South Africa
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Staffan Widstrand / African elephant (Loxodonta africana) male tusker walking down road photographed from vehicle, iMfolozi National Park, South Africa
White rhinoceros (Cerathorium simum) watched by anti-poaching patrol of iMfolozi National Park, South Africa October 2011
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Staffan Widstrand / White rhinoceros (Cerathorium simum) watched by anti-poaching patrol of iMfolozi National Park, South Africa October 2011
Common Zebra (Equus quagga) head detail portrait, Kruger National Park, South Africa
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Neil Aldridge / Common Zebra (Equus quagga) head detail portrait, Kruger National Park, South Africa
Portrait of a South African Parks Ranger whilst supervising a walking safari, Kruger National Park, Transvaal, South Africa, September 2008. No release available.
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Oriol Alamany / Portrait of a South African Parks Ranger whilst supervising a walking safari, Kruger National Park, Transvaal, South Africa, September 2008. No release available.
View of the night sky looking up through trees, with stars and the Milky Way, Mopani, Kruger National Park, Transvaal, South Africa, September.
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Oriol Alamany / View of the night sky looking up through trees, with stars and the Milky Way, Mopani, Kruger National Park, Transvaal, South Africa, September.
Close up of skin on the ear and back of an African elephant (Loxodonta africana), Mopani, Kruger National Park, Transvaal, South Africa, September.
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Oriol Alamany / Close up of skin on the ear and back of an African elephant (Loxodonta africana), Mopani, Kruger National Park, Transvaal, South Africa, September.
African elephant (Loxodonta africana) rear view Kruger National Park, Transvaal, South Africa, September.
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Oriol Alamany / African elephant (Loxodonta africana) rear view Kruger National Park, Transvaal, South Africa, September.
African Elephant (Loxodonta africana) browsing, Kruger National Park, Transvaal, South Africa, September.
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Oriol Alamany / African Elephant (Loxodonta africana) browsing, Kruger National Park, Transvaal, South Africa, September.
Male African lion (Panthera leo), Kruger National Park, Transvaal, South Africa, September.
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Oriol Alamany / Male African lion (Panthera leo), Kruger National Park, Transvaal, South Africa, September.
Cape Cobra (Naja nivea) female snake emerging from burrow. The snake often lives in abandoned mammal nests. deHoop Nature Reserve, Western Cape, South Africa.
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Tony Phelps / Cape Cobra (Naja nivea) female snake emerging from burrow. The snake often lives in abandoned mammal nests. deHoop Nature Reserve, Western Cape, South Africa.
Cape Hare (Lepus capensis) portrait. South Africa, December.
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Richard Du Toit / Cape Hare (Lepus capensis) portrait. South Africa, December.
Springbok (Antidorcas marsupialis) silhouetted on the horizon. Namaqualand, South Africa, December.
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Richard Du Toit / Springbok (Antidorcas marsupialis) silhouetted on the horizon. Namaqualand, South Africa, December.
Giraffes (Giraffa camelopardalis), two adults with baby, Kgalagadi Transfrontier Park, Northern Cape, South Africa, January
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Ann & Steve Toon / Giraffes (Giraffa camelopardalis), two adults with baby, Kgalagadi Transfrontier Park, Northern Cape, South Africa, January
Yellow mongoose (Cynictis penicillata) adult with young, Kgalagadi Transfrontier Park, South Africa, January
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Ann & Steve Toon / Yellow mongoose (Cynictis penicillata) adult with young, Kgalagadi Transfrontier Park, South Africa, January
Black backed jackal (Canis mesomelas) chasing doves from waterhole, Kgalagadi Transfrontier Park, South Africa, January 2012
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Ann & Steve Toon / Black backed jackal (Canis mesomelas) chasing doves from waterhole, Kgalagadi Transfrontier Park, South Africa, January 2012
Milne-edward's sifaka lemur (Propithecus diadema edwardsi) infant head portrait, Ranomafana National Park, south east Madagascar.
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Nick Garbutt / Milne-edward's sifaka lemur (Propithecus diadema edwardsi) infant head portrait, Ranomafana National Park, south east Madagascar.
Adult bull African Elephant (Loxononta africana) trunk reaching up to feed on foliage on the banks of the Luangwa River, South Luangwa National Park, Zambia, October
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Nick Garbutt / Adult bull African Elephant (Loxononta africana) trunk reaching up to feed on foliage on the banks of the Luangwa River, South Luangwa National Park, Zambia, October
Adult bull African Elephant (Loxononta africana) reaching up to feed on foliage on the banks of the Luangwa River, South Luangwa National Park, Zambia, October
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Nick Garbutt / Adult bull African Elephant (Loxononta africana) reaching up to feed on foliage on the banks of the Luangwa River, South Luangwa National Park, Zambia, October
Adult male Hippopotamus (Hippopotamus amphibius) posturing in agressive 'yawn' behaviour, Luangwa River, South Luangwa National Park, Zambia, November
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Nick Garbutt / Adult male Hippopotamus (Hippopotamus amphibius) posturing in agressive 'yawn' behaviour, Luangwa River, South Luangwa National Park, Zambia, November
Adult male Hippopotamuses (Hippopotamus amphibius) posturing in aggressive 'yawn' behaviour, Luangwa River, South Luangwa National Park, Zambia, November. Digital composite (two frames stitched togeth...
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Nick Garbutt / Adult male Hippopotamuses (Hippopotamus amphibius) posturing in aggressive 'yawn' behaviour, Luangwa River, South Luangwa National Park, Zambia, November. Digital composite (two frames stitched togeth...
Juvenile African elephant (Loxodonta africana) wanders through a tourist safari lodge, Mfuwe Lodge, South Luangwa NP, Zambia, October
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Nick Garbutt / Juvenile African elephant (Loxodonta africana) wanders through a tourist safari lodge, Mfuwe Lodge, South Luangwa NP, Zambia, October
Adult bull African Elephant (Loxononta africana) reaching up to feed on foliage on the banks of the Luangwa River, South Luangwa National Park, Zambia, October
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Nick Garbutt / Adult bull African Elephant (Loxononta africana) reaching up to feed on foliage on the banks of the Luangwa River, South Luangwa National Park, Zambia, October
African Elephant (Loxodonta africana) baby standing by its mother's legs. Kruger National Park, South Africa, December.
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Pete Oxford / African Elephant (Loxodonta africana) baby standing by its mother's legs. Kruger National Park, South Africa, December.
Baby African elephants (Loxodonta africana) of various ages playing together, Addo national park, Eastern Cape, South Africa, January. Did you know? Elephants can tell the difference between different...
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Ann & Steve Toon / Baby African elephants (Loxodonta africana) of various ages playing together, Addo national park, Eastern Cape, South Africa, January. Did you know? Elephants can tell the difference between different...
Green-grooved Dung Beetle (Scarabaeus rugosus) male with dung ball. deHoop NR, Western Cape, South Africa
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Tony Phelps / Green-grooved Dung Beetle (Scarabaeus rugosus) male with dung ball. deHoop NR, Western Cape, South Africa
Three Cape Ground Squirrels (Xerus inauris)  standing together at sunset. Kgalagadi TB Park of South Africa
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Charlie Summers / Three Cape Ground Squirrels (Xerus inauris) standing together at sunset. Kgalagadi TB Park of South Africa
Double rainbow over Kalahari, Kgalagadi Transfrontier Park, Botswana
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Vincent Grafhorst / Double rainbow over Kalahari, Kgalagadi Transfrontier Park, Botswana
Giant Pachypodium species photographed from the inside. Lake Tsimanampetsotsa National Park, South West Madagascar.
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Nick Garbutt / Giant Pachypodium species photographed from the inside. Lake Tsimanampetsotsa National Park, South West Madagascar.
Bottle Baobab (Adansonia rubrostipa) over 3000 years old. Lake Tsimanampetsotsa National Park, South West Madagascar.
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Nick Garbutt / Bottle Baobab (Adansonia rubrostipa) over 3000 years old. Lake Tsimanampetsotsa National Park, South West Madagascar.
Cheetah cub (Acinonyx jubatus) on the back of  juvenile Springbok (Antidorcas marsupialis)  Kgalagadi Trans Border Park, South Africa
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Charlie Summers / Cheetah cub (Acinonyx jubatus) on the back of juvenile Springbok (Antidorcas marsupialis) Kgalagadi Trans Border Park, South Africa
Meerkat / Suricate (Suricata suricatta) pregnant female standing on hind legs, with swollen abdomen exposed, Kgalagadi Transfrontier Park, South Africa
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Ann & Steve Toon / Meerkat / Suricate (Suricata suricatta) pregnant female standing on hind legs, with swollen abdomen exposed, Kgalagadi Transfrontier Park, South Africa
Cape fox cub (Vulpes chama) stretching,  Kgalagadi Transfrontier Park, Northern Cape, South Africa
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Ann & Steve Toon / Cape fox cub (Vulpes chama) stretching, Kgalagadi Transfrontier Park, Northern Cape, South Africa
Cape fox cub (Vulpes chama) carrying small rodent prey in mouth, Kgalagadi Transfrontier Park, Northern Cape, South Africa
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Ann & Steve Toon / Cape fox cub (Vulpes chama) carrying small rodent prey in mouth, Kgalagadi Transfrontier Park, Northern Cape, South Africa
Cape fox cubs (Vulpes chama) play-fighting, Kgalagadi Transfrontier Park, Northern Cape, South Africa
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Ann & Steve Toon / Cape fox cubs (Vulpes chama) play-fighting, Kgalagadi Transfrontier Park, Northern Cape, South Africa
Cape fox cubs (Vulpes chama) play-fighting, Kgalagadi Transfrontier Park, Northern Cape, South Africa
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Ann & Steve Toon / Cape fox cubs (Vulpes chama) play-fighting, Kgalagadi Transfrontier Park, Northern Cape, South Africa
Cape fox (Vulpes chama) cub licking its mothers face, Kgalagadi Transfrontier Park, Northern Cape, South Africa
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Ann & Steve Toon / Cape fox (Vulpes chama) cub licking its mothers face, Kgalagadi Transfrontier Park, Northern Cape, South Africa
Cape fox (Vulpes chama) sitting with cub, Kgalagadi Transfrontier Park, Northern Cape, South Africa
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Ann & Steve Toon / Cape fox (Vulpes chama) sitting with cub, Kgalagadi Transfrontier Park, Northern Cape, South Africa
Ground squirrel (Xerus inuaris) grooming baby, Kgalagadi Transfrontier Park, Northern Cape, South Africa Non-ex.
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Ann & Steve Toon / Ground squirrel (Xerus inuaris) grooming baby, Kgalagadi Transfrontier Park, Northern Cape, South Africa Non-ex.
Cape buffalo (Syncerus caffer caffer) displaying flehmen posture, to detect scent, Kruger national park, South Africa
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Ann & Steve Toon / Cape buffalo (Syncerus caffer caffer) displaying flehmen posture, to detect scent, Kruger national park, South Africa
Nile crocodile (Crocodylus niloticus) with jaws wide open, Kruger National Park, South Africa
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Ann & Steve Toon / Nile crocodile (Crocodylus niloticus) with jaws wide open, Kruger National Park, South Africa
Cape Ground Squirrel (Xerus inauris), Kgalagadi Transfrontier Park, Botswana
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Vincent Grafhorst / Cape Ground Squirrel (Xerus inauris), Kgalagadi Transfrontier Park, Botswana
The Ampitheatre, Royal Natal National Park, Drakensberg Mountains, KwaZuluNatal, South Africa
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David Noton / The Ampitheatre, Royal Natal National Park, Drakensberg Mountains, KwaZuluNatal, South Africa
Common zebra (Equus quagga) in the bush, Kruger National Park, South Africa
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David Noton / Common zebra (Equus quagga) in the bush, Kruger National Park, South Africa
Six young Impala {Aepyceros melampus} running down road in Kruger NP, South Africa
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Patricio Robles Gil / Six young Impala {Aepyceros melampus} running down road in Kruger NP, South Africa
Burchell's Zebra (Equus burchellii) males sparring, Itala Game Reserve, South Africa
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Richard Du Toit / Burchell's Zebra (Equus burchellii) males sparring, Itala Game Reserve, South Africa
RF- Black rhinoceros (Diceros bicornis) looking threatening, Etosha National Park, Namibia, June. Endangered species. (This image may be licensed either as rights managed or royalty free.)
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Tony Heald / RF- Black rhinoceros (Diceros bicornis) looking threatening, Etosha National Park, Namibia, June. Endangered species. (This image may be licensed either as rights managed or royalty free.)
RF- African elephant (Loxodonta africana) young males play fighting, Etosha National Park, Namibia, June. Endangered species. (This image may be licensed either as rights managed or royalty free.)
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Tony Heald / RF- African elephant (Loxodonta africana) young males play fighting, Etosha National Park, Namibia, June. Endangered species. (This image may be licensed either as rights managed or royalty free.)
African lion (Panthera leo) hunting Buffalo, South Luangwa, Zambia (non-ex)
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Andy Rouse / African lion (Panthera leo) hunting Buffalo, South Luangwa, Zambia (non-ex)
African elephant (Loxodonta africana) spraying water from trunk, bathing, Marakele NP, South Africa
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Andy Rouse / African elephant (Loxodonta africana) spraying water from trunk, bathing, Marakele NP, South Africa
African elephant (Loxidonta africana) female and calf in breeding herd, South Luangwa NP, Zambia
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Andy Rouse / African elephant (Loxidonta africana) female and calf in breeding herd, South Luangwa NP, Zambia
African elephant (Loxidonta africana) female and calf in breeding herd, South Luangwa NP, Zambia
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Andy Rouse / African elephant (Loxidonta africana) female and calf in breeding herd, South Luangwa NP, Zambia
African elephant (Loxodonta africana) very old bull (75+ years), Marakele NP, South Africa
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Andy Rouse / African elephant (Loxodonta africana) very old bull (75+ years), Marakele NP, South Africa
Bark detail of Quiver tree (Aloe dichotoma), Augrabies NP, Kalahari desert, South Africa
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Jouan & Rius / Bark detail of Quiver tree (Aloe dichotoma), Augrabies NP, Kalahari desert, South Africa
Cape ground squirrel (Xerus inauris) at burrow entrance, Kgalagadi Transfrontier Park, Kalahari desert, South Africa
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Jouan & Rius / Cape ground squirrel (Xerus inauris) at burrow entrance, Kgalagadi Transfrontier Park, Kalahari desert, South Africa
Tsamma melon (Citrullus lanatus) growing on the sand dune, Kgalagadi Transfrontier Park, Kalahari desert, South Africa
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Jouan & Rius / Tsamma melon (Citrullus lanatus) growing on the sand dune, Kgalagadi Transfrontier Park, Kalahari desert, South Africa
African wild cat (Felis sylvestris libyca), Kgalagadi Transfrontier Park, Kalahari desert, South Africa
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Jouan & Rius / African wild cat (Felis sylvestris libyca), Kgalagadi Transfrontier Park, Kalahari desert, South Africa
Cape ground squirrel (Xerus inauris) feeding on Gemsbok cucumber, Kgalagadi Transfrontier Park, Kalahari desert, South Africa
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Jouan & Rius / Cape ground squirrel (Xerus inauris) feeding on Gemsbok cucumber, Kgalagadi Transfrontier Park, Kalahari desert, South Africa
Yellow mongoose (Cynictis penicillata) standing on hind legs, Kgalagadi Transfrontier Park, Kalahari desert, South Africa
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Jouan & Rius / Yellow mongoose (Cynictis penicillata) standing on hind legs, Kgalagadi Transfrontier Park, Kalahari desert, South Africa
Male Red hartebeest (Alcelaphus buselaphus) fighting, Kgalagadi Transfrontier Park, Kalahari desert, South Africa
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Jouan & Rius / Male Red hartebeest (Alcelaphus buselaphus) fighting, Kgalagadi Transfrontier Park, Kalahari desert, South Africa
Cape ground squirrel (Xerus inauris) feeding on flowers during the rainy season, Kgalagadi Transfrontier Park, Kalahari desert, South Africa
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Jouan & Rius / Cape ground squirrel (Xerus inauris) feeding on flowers during the rainy season, Kgalagadi Transfrontier Park, Kalahari desert, South Africa
Two young Cape ground squirrels (Xerus inauris) at burrow entrance, Kgalagadi Transfrontier Park, Kalahari desert, South Africa
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Jouan & Rius / Two young Cape ground squirrels (Xerus inauris) at burrow entrance, Kgalagadi Transfrontier Park, Kalahari desert, South Africa
Cape ground squirrel (Xerus inauris) family at burrow entrance, Kgalagadi Transfrontier Park, Kalahari desert, South Africa
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Jouan & Rius / Cape ground squirrel (Xerus inauris) family at burrow entrance, Kgalagadi Transfrontier Park, Kalahari desert, South Africa
Black-Backed Jackal (Canis mesomelas) portrait, Kgalagadi Transfrontier Park, Kalahari desert, South Africa
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Jouan & Rius / Black-Backed Jackal (Canis mesomelas) portrait, Kgalagadi Transfrontier Park, Kalahari desert, South Africa
Hammerhead stork {Scopus umbretta} beside the Sabie river, Kruger NP, South Africa
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Richard Du Toit / Hammerhead stork {Scopus umbretta} beside the Sabie river, Kruger NP, South Africa
Burchell's Zebra (Equus burchellii) close-up of face of braying animal, Hluhluwe-umfolozi Game Reserve, South Africa
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Richard Du Toit / Burchell's Zebra (Equus burchellii) close-up of face of braying animal, Hluhluwe-umfolozi Game Reserve, South Africa
South African Giraffe (Giraffa giraffa giraffa) mother and young, Londolozi, Sabi-Sand Game Reserve, South Africa
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Richard Du Toit / South African Giraffe (Giraffa giraffa giraffa) mother and young, Londolozi, Sabi-Sand Game Reserve, South Africa
Gemsbok (Oryx gazella) running, Namib-Naukluft National Park, Namibia
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Richard Du Toit / Gemsbok (Oryx gazella) running, Namib-Naukluft National Park, Namibia
Hippopotamus in Luangwa river at sunrise, South Luangwa National Park, Zambia
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TJ Rich / Hippopotamus in Luangwa river at sunrise, South Luangwa National Park, Zambia
RF- Two Common zebra (Equus quagga) fighting, Etosha national park, Namibia. (This image may be licensed either as rights managed or royalty free.)
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Tony Heald / RF- Two Common zebra (Equus quagga) fighting, Etosha national park, Namibia. (This image may be licensed either as rights managed or royalty free.)
Bat Eared Fox emerging from den (Otocyon megalotis) Kalahari Gemsbok NP South Africa
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Tony Heald / Bat Eared Fox emerging from den (Otocyon megalotis) Kalahari Gemsbok NP South Africa
Pied kingfisher {Ceryle rudis} hovering, Kruger National Park, South Africa.
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Hanne & Jens Eriksen / Pied kingfisher {Ceryle rudis} hovering, Kruger National Park, South Africa.
RF- Common zebra foal running (Equus quagga). Etosha National Park, Namibia. Digitally enhanced. (This image may be licensed either as rights managed or royalty free.)
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Tony Heald / RF- Common zebra foal running (Equus quagga). Etosha National Park, Namibia. Digitally enhanced. (This image may be licensed either as rights managed or royalty free.)
Yellow Baboon {Papio cynocephalus} Young riding on mother's back, Kruger National Park, South Africa
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Dave Watts / Yellow Baboon {Papio cynocephalus} Young riding on mother's back, Kruger National Park, South Africa
Hippopotamus at water surface. Kruger NP, South Africa
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Tony Heald / Hippopotamus at water surface. Kruger NP, South Africa
Cheetah (Acinonyx jubatus) adult female laying in the grass with her three month old cubs, summer, Phinda Game Reserve, South Africa
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Richard Du Toit / Cheetah (Acinonyx jubatus) adult female laying in the grass with her three month old cubs, summer, Phinda Game Reserve, South Africa
South African Giraffe (Giraffa giraffa giraffa) mother grooming her calf, Phinda Game Reserve, South Africa
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Richard Du Toit / South African Giraffe (Giraffa giraffa giraffa) mother grooming her calf, Phinda Game Reserve, South Africa
Springbok (Antidorcas marsupialis) grazing under tree, Kalahari Gemsbok Park, South Africa
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Richard Du Toit / Springbok (Antidorcas marsupialis) grazing under tree, Kalahari Gemsbok Park, South Africa
Red-billed Oxpecker (Buphagus erythrorhynchus) on Impala (Aepyceros melampus) male, Kruger National Park, South Africa
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Richard Du Toit / Red-billed Oxpecker (Buphagus erythrorhynchus) on Impala (Aepyceros melampus) male, Kruger National Park, South Africa
African Scarab Beetle (Circellium bacchus) rolling dung ball, Addo National Park, South Africa
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Mark Moffett / African Scarab Beetle (Circellium bacchus) rolling dung ball, Addo National Park, South Africa
Praying Mantis (Mantis sp), Ndumo Game Reserve, South Africa
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Gerry Ellis / Praying Mantis (Mantis sp), Ndumo Game Reserve, South Africa
Ant (Formicidae) pair communicating, Karoo National Park, western South Africa
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Gerry Ellis / Ant (Formicidae) pair communicating, Karoo National Park, western South Africa
African wild dogs {Lycaon pictus} playfighting. Savute, Chobe NP, Botswana.
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Richard Du Toit / African wild dogs {Lycaon pictus} playfighting. Savute, Chobe NP, Botswana.
View of Tugela river and Amphitheatre, Royal Natal NP, KZN, South Africa
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Pat De La Harpe / View of Tugela river and Amphitheatre, Royal Natal NP, KZN, South Africa
Gemsbok in long grass {Oryx gazella gazella} Kruger NP South Africa semi desert habitat
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Richard Du Toit / Gemsbok in long grass {Oryx gazella gazella} Kruger NP South Africa semi desert habitat
Cheetah chasing Jackal. (Acininxy jubatus) South Africa Kalahari Gemsbok NP.
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Tony Heald / Cheetah chasing Jackal. (Acininxy jubatus) South Africa Kalahari Gemsbok NP.
Hippopotamus submerged, eyes and ears just above water.Kruger National Park, South Africa
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Tony Heald / Hippopotamus submerged, eyes and ears just above water.Kruger National Park, South Africa
Male Impala running in Kruger National Park, South Africa. Impala can run at speeds of 50mph!
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Richard Du Toit / Male Impala running in Kruger National Park, South Africa. Impala can run at speeds of 50mph!