Steve Knell / Song thrush (Turdus philomelos clarkei) portrait, Hungary, May.

Steve Knell / Bearded tit (Panurus biarmicus) male in summer plumage. Hungary, May

Steve Knell / Whooper swan (Cygnus cygnus) on water, backlit showing swarming midges. Iceland, May

Steve Knell / Harlequin duck (Histrionicus histrionicus) portrait of male stood motionless on exposed rock, in river, Iceland, May

Steve Knell / Harlequin ducks (Histrionicus histrionicus) male chasing female on riverbank, Iceland, May

Steve Knell / Tufted duck (Aythya fuligula) drake on water with Midges (Chironomidae) Iceland. June

Steve Knell / Boreray, under low cloud cover. This holds the largest gannetry in UK (60,000 pairs) St. Kilda, Scotland, July 2009

Steve Knell / Long-tailed duck (Clangula hyemalis) male in summer plumage, taking off from river, Iceland, May

Steve Knell / Harlequin duck (Histrionicus histrionicus) male standing in rapids, Iceland, May

Steve Knell / Swarm of Midges (Chironomidae) flying around grass stems. Lake Myvatn, Iceland. June

Steve Knell / Swarm of Midges (Chironomidae) flying around grass stems. Lake Myvatn, Iceland. June

Steve Knell / Common starling (Sturnus vulgaris) flocking prior to roosting, Scotland, January 2009