Martin Dohrn / Herd of giraffes (Giraffa camelopardalis) resting at night, Masai Mara, Kenya. Image taken at night using thermal camera technology, with no artificial light.

Will Burrard-Lucas / White rhino (Ceratotherium simum), portrait. Solio Game Reserve, Solio Ranch, Kenya. Taken with remote camera buggy / BeetleCam. EDITORIAL USE ONLY. All other uses require clearance

Shane Gross / Invasive Lionfish (Pterois volitans) which have taken over and are wiping out native fish in the Atlantic ocean. The highest densities are in the northern gulf of Mexico. Destin, Florida, USA.

Shane Gross / Invasive Lionfish (Pterois volitans) which have taken over and are wiping out native fish in the Atlantic ocean. The highest densities are in the northern gulf of Mexico. Destin, Florida, USA.

David Gallan / Two Southern right whales (Eubalaena australis) playing in shallows, taken with drone, Tathra Beach, New South Wales, Australia, August.

Sven Zacek / Ural owl (Strix uralensis) female in flight having taken off from nest, in evening light. Tartumaa, Southern Estonia. May.

Will Burrard-Lucas / Black and white image of African elephant (Loxodonta africana) Tsavo Conservation Area, Kenya. Taken with a remote camera buggy / BeetleCam. Editorial use only.

Will Burrard-Lucas / Black and white image of African elephant (Loxodonta africana) herd with calf at waterhole, seen through adults legs. Tsavo Conservation Area, Kenya. Taken with a remote camera buggy / BeetleCam. Edit...

Terry Whittaker / Red fox (Vulpes vulpes) adult with train in background, Kent, UK. March. Highly commended in the Nature Photographer of the Year Awards 2018. Taken with remote camera.

Ralph Pace / Humpback Whale (Megaptera novaeangliae) male and female courting, Maui, Hawaii, image taken under NMFS Permit # 19225

Jen Guyton / Park ranger holding a Cape pangolin / Temminck's ground pangolin (Smutsia temminckii), rescued from poachers. This picture was taken shortly before freeing the pangolin. Gorongosa National Park, Moza...

Theo Bosboom / Autumn leaves from Beech, Oak and Birch trees floating on water, taken from underwater, Ardennes, La Hoegne, Belgium.

Ingo Arndt / Grizzly bear (Ursus arctos horribilis) picture taken with a remote camera car, Alaska, July.

Ingo Arndt / African wild dog portrait {Lycaon pictus} Namibia. Taken under controlled conditions.

Neil Aldridge / White rhinoceros (Ceratotherium simum) female 'Thandi' silhouetted against the sunset, Kariega Game Reserve. South Africa. Thandi's horns were taken in a brutal attack by poachers.

Will Burrard-Lucas / Lion (Panthera leo) walking over the plain, taken with remote camera. Serengeti National Park, Tanzania. December

Will Burrard-Lucas / Spotted hyena (Crocuta crocuta) at night, taken with remote camera. Liuwa Plain National Park, Zambia. December

Will Burrard-Lucas / Spotted hyena (Crocuta crocuta) low angle shot of group taken with remote camera. Liuwa Plain National Park, Zambia. May

Anup Shah / African elephant (Loxodonta africana) approaching camera with curiosity, Maasai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / Lion (Panthera leo) male approaching, Maasai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / African elephants (Loxodonta africana) feeding on loose soil for its minerals, Maasai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / African elephants (Loxodonta africana) family herd feeding on loose soil for its minerals, with dramatic stormy skies behind, Maasai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / African elephants (Loxodonta africana) family herd feeding on loose soil for its minerals, with dramatic stormy skies behind, Maasai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Alex Mustard / Boulder brain coral (Colpophyllia natans) photographed with a long exposure with camera rotation. Taken with circular fisheye lens. Grand Cayman, Cayman Islands. Caribbean Sea.

Tony Wu / Humpback whale (Megaptera novaeangliae) rear view, taken while the whale was staying still and singing. Hunga Channel. Vava'u, Kingdom of Tonga, Pacific Ocean.

Wim van den Heever / Aerial view of Cape Town city with Table Mountain, South Africa, taken from helicopter, May 2011

Wim van den Heever / Aerial view of Cape Town city with Table Mountain, South Africa, taken from helicopter, May 2011

Wim van den Heever / Aerial view of Cape Town city with Table Mountain, South Africa, taken from helicopter, May 2011

Wim van den Heever / African Wild dogs or Cape hunting dogs (Lycaon pictus) at close range taken from ground level, Zimanga Private Game Reserve, South Africa.

Charlie Summers / Andrommeda (M31) with its two syster galaxies (M32 and M110) Eastern Colorado. 8 November 2015. Taken with digital focus stacking.

Wim van den Heever / Black rhinoceros (Diceros bicornis) mother and calf having a drink at night with perfect reflections. Etosha National Park, Namibia. Taken with infrared camera.

Wim van den Heever / Black rhinoceros (Diceros bicornis) bull having a drink at night with perfect reflections. Etosha National Park, Namibia. Taken with Infrared camera.

Eric Baccega / African elephant (Loxodonta africana) and black rhinoceros (Diceros bicornis) bathing at waterhole at night with moon, Okaukuejo, Etosha National Park, Namibia, Africa. Taken with infrared camera.

Ben Hall / Marine iguana (Amblyrhynchus cristatus) on rock taken with slow shutter speed to show motion in the water, Galapagos islands, May.

Wim van den Heever / Nile crocodile (Crocodylus niloticus) still in the fast flowing water of the Mara River, taken with slow shutter speed. Masai Mara, Kenya.

Alex Hyde / View up towards autumnal Beech (Fagus sylvatica) woodland canopy, taken with a fisheye lens. Derbyshire, UK. October.

Staffan Widstrand / Bonelli's eagle or Eurasian hawk-eagle, Hieraetus fasciatus or Aquila fasciata, picture taken from hide, at a feeding station for conservation purposes, utillizing live domestic pigeons caught as pest...

Gerrit Vyn / Snowy Owl (Bubo scandiacus) fluffing feathers, this image was taken during a massive increase in the range and population of snowy owls. Boundary Bay, Canada. March.

Claudio Contreras / Caribbean flamingo (Phoenicopterus ruber) about to take off to roosting site, Ria Celestun Biosphere Reserve, Yucatan Peninsula Mexico, January. Best image taken in Mexico and specially commended in...

Bence Mate / Wild boar (Sus scrofa) at night with the milky way in the background, Gyulaj, Tolna, Hungary. August. Taken using long exposure with flash at night. Winner of the Mammals Category of the GDT Awards 20...

Eric Medard / Roe deer (Capreolus capreolus) female, taken at night using infra-red, Mayenne, Pays de la Loire, France.

Eric Medard / Barn owl (Tyto alba) flying in old chapel at night, taken with infra red remote camera trap, Mayenne, Pays de Loire, France, November.

Alex Mustard / Nudibranch (Flabellina lineata) in aquarium, taken in the field in Gulen, Norway. North East Atlantic Ocean. Collected for scientific research.

Pete Oxford / Konyak Naga tattooed head hunter. Only someone who has killed and taken a human head can be tattooed in this manner. Mon district. Nagaland, North East India, October 2014.

Pete Oxford / Konyak Naga tattooed Ang or Chief. Only someone who has killed and taken a human head can be tattooed in this manner. Mon district. Nagaland, North East India, October 2014.

Pete Oxford / Konyak Naga tattooed Ang or Chief. Only someone who has killed and taken a human head can be tattooed in this manner. Mon district. Nagaland, North East India, October 2014.

Bruno D'Amicis / Fennec fox (Vulpes zerda) adult at night digging for prey among the roots of Retam broom shrub. Grand Erg Oriental, Kebili Governorate, Tunisia. Taken with remote camera trap.

Bruno D'Amicis / Fennec fox (Vulpes zerda) foraging at night, Grand Erg Oriental, Kebili Governorate. Tunisia. Taken with remote camera trap.

Bruno D'Amicis / Fennec fox (Vulpes zerda) displayed in the market (souk), of a Tunisian town. This species is adapted to silence and solitude of the desert, this individual that had been caught as an adult showed cle...

Bruno D'Amicis / Fennec fox (Vulpes zerda) displayed in the market (souk), of a Tunisian town. This species is adapted to silence and solitude of the desert, this individual that had been caught as an adult showed cle...

Nick Upton / Eurasian beaver (Castor fiber) gnawing the trunk of a Willow tree (Salix sp.) in woodland enclosure at night. Devon Beaver Project, run by Devon Wildlife Trust, Devon, UK, April. Taken by a remote cam...

Nick Upton / Eurasian beaver (Castor fiber) entering the River Otter after release. Part of escaped population re-released to the wild following veterinary check ups. Taken on the second day of Beaver releases aft...

Nick Upton / Julia Coats of the Animal and Plant Health Agency releasing Eurasian beaver (Castor fiber) back into River Otter. Part of escaped population re-released to the wild following veterinary check ups. Tak...

Charlie Summers / The Pleiades or Seven Sisters (M45), a star cluster in the constellation Taurus, taken 26 September 2014 from Eastern Colorado. Taken with digital focus stacking.

Gavin Hellier / Timelapse taken from the back of a rickshaw travelling through Bangkok, Thailand, 2011.

Gavin Hellier / Timelapse of tourists having their photographs taken in front of the Taj Mahal, Taj Mahal UNESCO World Heritage Site, Agra, Uttar Pradesh, India, 2011.

Ammonite / Baird's tapir (Tapirus bairdii) entering a pool of water and wallowing, footage taken at night using thermal camera technology, Santa Rosa National Park, Costa Rica.

Ammonite / Group of African lion (Panthera leo) cubs playing with their mother and another lioness, footage taken at night using starlight camera technology, Masai Mara, Kenya.

Ammonite / Group of African lion (Panthera leo) cubs play fighting with each other, footage taken at night using starlight camera technology, Masai Mara, Kenya.

Ammonite / Group of African lion (Panthera leo) cubs play fighting with each other, footage taken at night using starlight camera technology, Masai Mara, Kenya.

Ammonite / African lion (Panthera leo) cub playing with mother before walking away, , footage taken at night using starlight camera technology, Masai Mara, Kenya.

Ammonite / African lioness (Panthera leo) growling and swatting at a sub-adult male approaching a kill, footage taken at night using thermal camera technology, Masai Mara, Kenya.

Ammonite / Group of sub-adult African lions (Panthera leo) chasing and jumping on a mother Hippopotamus (Hippopotamus amphibius) and her calf, footage taken at night using infrared camera technology, Masai Mara,...

Ammonite / Sub-adult African lioness (Panthera leo) chasing and jumping onto the back of a Hippopotamus (Hippopotamus amphibius), footage taken at night using infrared camera technology, Masai Mara, Kenya.

Ammonite / Large group of Black wildebeest (Connochaetes gnou) migrating across savanna in a line, footage taken at night using thermal camera technology, Masai Mara, Kenya.

Ammonite / Close-up of a Yellow slug (Limax flavus) moving towards the camera leaving a slime trail and extending and retracting its eyestalk, footage taken using an endoscopic camera, Bristol, England, UK, Marc...

Ammonite / Earthworm (Lumbricus terrestris) moving through leaf litter, footage taken using an endoscopic camera in controlled conditions, Bristol, England, UK, June.

Ammonite / Close-up of a Yellow slug (Limax flavus) leaving a slime trail as it moves, footage taken using an endoscopic camera, Bristol, England, UK, March.

Ammonite / Water buffalo (Bubalus arnee) chasing Sri Lankan leopard (Panthera pardus kotiya) near a waterhole, footage taken at night using thermal camera technology, Yala National Park, Sri Lanka.

Ammonite / Family of Sri Lankan elephant (Elephas maximus maximus) drinking at waterhole, footage taken at night using thermal camera technology, showing cooling thermoregulatory properties of ears, Yala Nationa...

Alex Mustard / Portrait of Bandcheek wrasse (Oxycheilinus digrammus) taken during the day. Anemone City, Ras Mohammed Marine Park, Gulf of Aqaba, Red Sea. Sinai, Egypt.

Will Burrard-Lucas / Big-headed mole rat (Tachyoryctes macrocephalus) appearing from its hole, Bale Mountains National Park, Ethiopia. Taken with remote camera.

Bence Mate / Grey heron (Ardea cinerea) catching fish, taken with remote camera, Pusztaszer, Hungary, June.

Dong Lei / Blood pheasants (Ithaginis cruentus) foraging in habitat, one looking at the camera, Meli Snow Mountain National Park, Yunnan Province, China, January. Endemic. Taken with remote camera.

Jabruson / Vehicle carrying several different duiker carcasses for commercial bushmeat trade including: Blue Duiker (Cephalophus monticola), Bay Duiker (Cephalophus dorsalis), Peter's Duiker (Cephalophus callipy...

Anup Shah / Eastern white-bearded wildebeest (Connochaetes taurinus) herd running. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / African elephants (Loxodonta africana) drinking at a waterhole. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / African elephants (Loxodonta africana) wallowing in a waterhole. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / African elephants (Loxodonta africana) dust bathing. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / African elephants (Loxodonta africana) at a waterhole. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / African elephant (Loxodonta africana) herd in the savanna. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / Black-backed jackals (Canis mesomelas) feeding on a wildebeest carcass. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / Black-backed jackal (Canis mesomelas) standing next to a wildebeest carcass. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / Spotted hyena (Crocuta crocuta) pup approaching with curiosity. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / Spotted hyenas (Crocuta crocuta) approaching with curiosity. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / Spotted hyena (Crocuta crocuta) approaching with curiosity. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / Olive baboon (Papio anubis) sitting portrait. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / Lion (Panthera leo) old male with one eye standing next to a kill. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / Common or Plains zebra herd crossing the Mara River - wide angle perspective (Equus quagga burchellii). Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / White-backed vultures (Gyps africanus) feeding on the carcass of a Common or Plains zebra. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / Eastern white-bearded wildebeest (Connochaetes taurinus) herd crossing the Mara River watched by tourists. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / Olive baboon (Papio anubis) juveniles watching with curiosity. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / Spotted hyena (Crocuta crocuta) approaching with curiosity. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / Spotted hyena (Crocuta crocuta) aged about 2 years watching with curiosity with the rest of the clan in the background. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / Lion (Panthera leo) cub aged about 2 years approaching with curiosity - remote wide angle perspective . Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / Eastern white-bearded wildebeest (Connochaetes taurinus) herd running. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.