Paul D Stewart / Chromolithograph illustration of Cromagnon 'Skull of an old man' Plate 1, Section C. Edouard Lartet and Henry Christy. 'Reliquiae Aquitanicae' 1865-1875, Williams and Norgate, London 1875..

Paul Williams / Giant otter (Pteronura brasiliensis) adult playing with plastic bottle, Pantanal, Pocone, Brazil

Paul Williams / Tourists watching two wild Jaguar cubs fight on riverbank whilst third looks on from above (Panthera onca) from boat on river, Pantanal, Pocone, Brazil. No release available.

Rod Williams / Northern / Chinese white-cheeked gibbon (Nomascus leucogenys) female head portrait, Laos, Vietnam, critically endangered species. Captive.

Rod Williams / Young male North Chinese Leopard (Panthera pardus japonensis). Captive. Endemic to northern China. Endangered. UK, June.

Rod Williams / North American Porcupine (Erethizon dorsatum). Captive. Endemic to North America. Captive. UK, May.

Alan Williams / Oystercatcher (Haematopus ostralegus) perched on rock on shore, North Uist, Outer Hebrides, Scotland, UK, May

Alan Williams / Two Red Kites (Milvus milvus) fighting in flight. Mid-Wales, UK, October 2010.

Rod Williams / Francois' Langur (Trachypithecus francoisi) captive, from Vietnam, Laos and Southern China, Endangered

Rod Williams / Male Red-backed Bearded saki monkey (Chiropotes chiropotes) captive, from Venezuela

Alan Williams / Male Ruddy Duck (Oxyura jamaicensis) displaying on lake expelling air from breast feathers to form bubbles (captive bird) UK, May 2010

Rod Williams / Green / Emerald Tree Monitor (Varanus prasinus) head portrait climbing tree trunk. Captive, from Papua New Guinea, Indonesia

Rod Williams / Peacock praying mantis (Pseudempusa pinnapavonis) female with colourful wings open in defensive display. Captive, found in Thailand. May

Rod Williams / Peacock praying mantis (Pseudempusa pinnapavonis) female in defensive display with colourful wings. Captive, found in Thailand.

Rod Williams / Yellow-banded Poison Dart Frog (Dendrobates leucomelas) sitting on colourful variegated leaves. Captive, found in Brazil, Guyana, Venezuela

Alan Williams / Black-tailed Godwits (Limosa limosa) and Knots (Calidris canutus) in flight in early morning light, Dee Estuary, Merseyside, England, UK, February

Alan Williams / Robin (Erithacus rubecula) perched in snow covered branches, Cheshire, UK, January

Alan Williams / Robin (Erithacus rubecula) on snow covered lawn, Cheshire, UK, January

Alan Williams / Dunnock (Prunella modularis) perched on fence in garden, Cheshire, UK, January

Alan Williams / Female blackcap (Sylvia atricapilla) perched in garden shrub, Cheshire, UK, December

Alan Williams / Grey Heron (Ardea cinerea) in flight, being chased by Lapwing (Vanellus vanellus) North Wales, UK, May

Alan Williams / Wood warbler (Phylloscopus sibilatrix) singing in woodland, North Wales, UK, May

Alan Williams / Short eared owl (Asio flammeus) on roadside post, Norfolk, UK, January

Alan Williams / Skylark (Alauda arvensis) singing while hovering, Cheshire, UK, April

Alan Williams / Yellow wagtail (Motacilla flava flavissima) male singing perched on Rapeseed flower head, Essex, UK, May

Alan Williams / Short eared owl (Asio flammeus) hovering while hunting in flight, Norfolk, UK, January

Alan Williams / Turnstone (Arenaria interpres) in flight at tide line in winter plumage with tail spread for landing, Whitstable Bay, Kent, UK, December

Alan Williams / Barn swallow (Hirundo rustica) in flight, Cheshire, UK, June

Alan Williams / Common teal (Anas crecca) male displaying, Martin Mere WWT, Lancashire, UK, February

Alan Williams / Bohemian waxwing (Bombycilla garrulus) perched in Rowan tree (Sorbus) feeding on berries, North Wales, UK, January

Alan Williams / Bohemian waxwing (Bombycilla garrulus) taking off from Rowan tree (Sorbus) with full crop after eating berries, North Wales, UK, January

Alan Williams / Stonechat (Saxicola torquata) male taking off from Gorse, Suffolk, UK, May

Alan Williams / Whitethroat (Sylvia communis) male singing perched on Blackthorn (Prunus spinosa) blossom, Essex, UK, May

Alan Williams / Bohemian waxwing (Bombycilla garrulus) perched in Rowan tree (Sorbus sp) North Wales, UK, January

Rod Williams / Scarlet macaws (Ara macao) in flight, captive, from Central and South America

Rod Williams / Golden Poison Dart Frog (Phyllobates terribilis) captive, from Colombia, Endangered Species

Alan Williams / Pintail (Anas acuta) male flapping wings after bathing, Martin Mere WWT reserve, Lancashire, UK, February

Alan Williams / European rabbit (Oryctolagus cuniculus) sitting up in meadow on alert, Essex, UK, June

Alan Williams / Ruff (Philomachus pugnax) male foraging along shore in winter plumage, Lancashire, UK, February

Alan Williams / Grey heron (Ardea cinerea) pair at nest, lower bird adding twig to nest, Midlands, UK, February

Alan Williams / Blue tit (Parus caeruleus) perched on twig with catkins, Essex, UK, February

Alan Williams / Blue tit (Parus caeruleus) calling, perched on blossom covered twig, Essex, UK, March

Alan Williams / Brent geese (Branta bernicla) flock flying over farmland, Whitstable, Kent, UK, January

Alan Williams / Male Bullfinch (Pyrrhula pyrrhula) feeding on Dock (Rumex) seeds, Yorkshire, UK, January

Alan Williams / Male Blackcap (Sylvia atricapilla) singing in woodland, Essex, UK, April

Rod Williams / Himalayan / Crocodile Newt (Tylotriton verrucosus) on mossy rock. Occurs SE Asia, NE India and E Nepal. June 2OO8, Captive.

Rod Williams / Dyeing poison arrow frog {Dendrobates tinctorius} sitting on leaf. Captive.

Rod Williams / Female Geoffroy's cat {Felis geoffroyi}. Rare melanistic form. Captive.

Rod Williams / Sloth bear {Melursus ursinus} female portrait. Captive. Sri Lanka.

Rod Williams / Malayan tapir {Tapirus indicus}, Thailand. Captive

Rod Williams / Ring-tailed lemur {Lemur catta} infant on adults back. Captive. Madagascar. Lemurs

Rod Williams / Lowland anoa {Bubalus depressicornis} grazing. Captive. Indonesia

Rod Williams / Aardwolf {Proteles cristatus} sub-adult female. Captive.

Rod Williams / Mexican red-knee tarantula {Brachypelma smithii}. Captive.

Rod Williams / Male Marbled cat {Felis marmorata} sitting on a log. Captive.

Rod Williams / Cotton-top tamarin {Saguinus oedipus} portrait. Captive, from Colombia.

Rod Williams / The Cutty Sark ship at Greenwich, London, UK.

Rod Williams / Two young Fishing cats {Prionailurus / Felis viverrinus} drinking. Captive.

Rod Williams / The Gerkin building in the city of London, UK.

Rod Williams / Two young Fishing cats {Prionailurus / Felis viverrinus} play-fighting. Captive.

Rod Williams / Barasingha / Swamp deer {Cervus duvauceli} female with fawn. Captive. India.

Rod Williams / Margay cat {Felis wiedii} male, captive.

Rod Williams / Black-footed cat {Felis nigripes} portrait. Captive.

Rod Williams / Celebes / Black / Sulawesi crested macaque {Macaca nigrai} female with infant. Captive.

Rod Williams / Sumatran orang utan {Pongo pygmaeus abelii} male portrait. Captive. Indonesia.

Rod Williams / Leaf insect {Phyllium giganteum} on leaf. Captive.

Rod Williams / Leadbeaters possum {Gymnobelideus leadbeateri} portrait, Australia. Captive

Rod Williams / Ratel / Honey badger {Mellivora capensis} captive, occurs in Africa, India & SW Asia, female

Rod Williams / Congo peafowl {Afropavo congensis} male. Captive.

Rod Williams / White-handed / Lar gibbon {Hylobates lar} male. Portrait showing teeth. Captive.

Rod Williams / Goeldis monkey {Callimico goeldii} lying on a branch. Captive from Brazil.

Rod Williams / Two Dorcas gazelles {Gazella dorcas neglecta} resting. Captive.

Rod Williams / Two Black-billed / West Indian whistling duck {Dendrocygna arborea} Cuba. Captive

Rod Williams / Fishing cat {Prionailurus / Felis viverrinus} male, captive.

Rod Williams / Male Geoffroy's cat {Felis geoffroyi} sitting on branch. Captive.

Rod Williams / Leadbeaters possum {Gymnobelideus leadbeateri} portrait, Australia. Captive

Rod Williams / Hermit / Waldrapp / Northen bald ibis {Geronticus eremita} portrait. Captive.

Rod Williams / Two Formosan sika deer {Cervus nippon taiouanus} males. Captive. Taiwan.

Rod Williams / Blue crane {Grus paradisea} portrait. Captive.

Rod Williams / Black-billed / West Indian whistling duck {Dendrocygna arborea}, Cuba. Captive

Rod Williams / Young male Aardwolf {Proteles cristatus} in long grass. Captive.

Rod Williams / The Royal Naval College, Greenwich, London, UK.

Rod Williams / Young Fishing cat {Prionailurus / Felis viverrinus} by water, captive

Rod Williams / Black mangabey, young male {Cercocebus / Lophocebus aterrimus} from central Africa - captive

Rod Williams / Black mangabey, young male {Cercocebus / Lophocebus aterrimus} from central Africa. Captive

Rod Williams / Crocodile newt {Tylototriton verrucosus} from Asia

Rod Williams / Giant asian praying mantis {Hierodula membranacea} male, captive from India

Rod Williams / Red Panda {Ailurus fulgens} resting in tree. Captive, occurs India, Nepal, China, Laos.

Rod Williams / Western gentle lemur {Hapalemur griseus occidentalis} captive occurrs Madagascar, vulnerable

Rod Williams / Night monkey {Aotus trivirgatus} juvenile, captive, from Central and South America

Rod Williams / Lac Alaotra bamboo lemur juvenile {Hapalemur griseus alaotrensis} Madagascar Lack

Rod Williams / Young Common marmosets {Callithrix jacchus} native to north-east Brazil, S America

Rod Williams / Male Chinese water deer {Hydropotes inermis} native to China

Rod Williams / Female Alaotran gentle lemur {Hapalemur griseus alaotrensis} Madagascar. Critically endangered species

Rod Williams / Young male Black Concolor gibbon portrait {Hylobates concolor} occurs Vietnam, N Laos, Isle of Hainan

Rod Williams / Portrait of Golden lion tamarin {Leontopithecus rosalia} occurs Brazil, Rio de Janeiro State