Nick Upton / Jerusalem thorn / Christ's thorn / Crown of thorns tree (Paliurus spina-christi) with clusters of yellow winged fruits / seed pods growing on coastal limestone hillside, Zadar province, Croatia, July.

Nick Upton / Overview of Vinjerac fishing village and harbour with Pine trees (Pinus sp.) in the foreground and the karst limestone Velebit mountain range of the Dinaric Alps in the background, Zadar province, Cro...

Nick Upton / Overview of Vinjerac fishing village and harbour with Pine trees (Pinus sop.) in the foreground and the karst limestone Velebit mountain range of the Dinaric Alps in the background, Zadar province, Cr...

Nick Upton / Overview of Vinjerac fishing village and harbour with Pine trees (Pinus sp.) in the foreground and the karst limestone Velebit mountain range of the Dinaric Alps in the background, Zadar province, Cro...

Nick Upton / Common golden thistle / Spanish oyster thistle (Scolymus hispanicus) with clusters of yellow flowers growing on coastal limestone hillside, Zadar province, Croatia, July.

Nick Upton / Common golden thistle / Spanish oyster thistle (Scolymus hispanicus) with clusters of yellow flowers growing on coastal limestone hillside, Zadar province, Croatia, July.

Nick Upton / Large Robber Fly (Machimus modestus) female with small solitary bee prey. Zadar province, Croatia, July.

Nick Upton / Large Robber Fly (Machimus modestus) male resting on rock. Zadar province, Croatia, July.

Nick Upton / Bee Fly (Hemipenthes velutina) feeding from Spiny Sow Thistle (Sonchus asper) flower. Zadar province, Croatia, July.

Nick Upton / Sailing yacht silhouetted at sea off Zadar as the sun sets, with Pasman island in the far background, Croatia, July.

Nick Upton / Keeled, winged seed pods of Jerusalem thorn / Christ's thorn / Crown of thorns tree (Paliurus spina-christi) against blue sky, growing on coastal limestone hillside, Zadar province, Croatia, July.

Nick Upton / Keeled, winged seed pods of Jerusalem thorn / Christ's thorn / Crown of thorns tree (Paliurus spina-christi) growing on coastal limestone hillside, Zadar province, Croatia, July.

Nick Upton / Chicory (Cichorium intybus) flower clump growing low to the ground on heavily sheep grazed coastal limestone hillside, Zadar province, Croatia, July

Nick Upton / Hermit Butterfly (Chazara briseis) feeding on a Slender Thistle / Italian Thistle (Carduus pycnocephalus) flower. Zadar province, Croatia, July.

Nick Upton / Egyptian Locust / Egyptian Grasshopper (Anacridium aegyptium) sunbasking on limestone rock. Zadar province, Croatia, July.

Nick Upton / Large Robber Fly (Machimus modestus) female with small solitary bee prey. Zadar province, Croatia, July.

Nick Upton / Shield Bug / Stink Bug (Carpocoris pudicus) on a thistle leaf. Zadar province, Croatia, July.

Nick Upton / Leafcutter Bee (Megachile sp.) feeding on Slender Thistle / Italian Thistle (Carduus pycnocephalus) flower. Zadar province, Croatia, July.

Nick Upton / Leafcutter Bee (Megachile sp.) feeding on Slender Thistle / Italian Thistle (Carduus pycnocephalus) flower. Zadar province, Croatia, July.

Nick Upton / Bee Fly (Anthrax leucogaster) sunbasking on limsetone rock. Zadar province, Croatia, July.

Nick Upton / Shield Bug / Stink Bug (Carpocoris pudicus) on a Spiny Sow Thistle (Sonchus asper) leaf. Zadar province, Croatia, July.

Nick Upton / Hermit Butterfly (Chazara briseis) resting on a Slender Thistle / Italian Thistle (Carduus pycnocephalus) flower. Zadar province, Croatia, July.

Nick Upton / Bee Fly (Geron sp.) with proboscis feeding from Slender Thistle / Italian Thistle (Carduus pycnocephalus) flower. Zadar province, Croatia, July.

Nick Upton / Bee Fly (Geron sp.) with proboscis feeding from Slender Thistle / Italian Thistle (Carduus pycnocephalus) flower. Zadar province, Croatia, July.

Nick Upton / Novigrad fishing village, Zadar-Knin region, Croatia with Velebit mountain range in background. 2007