Will Burrard-Lucas / White rhino (Ceratotherium simum), portrait. Solio Game Reserve, Solio Ranch, Kenya. Taken with remote camera buggy / BeetleCam. EDITORIAL USE ONLY. All other uses require clearance

Doug Perrine / Humpback whale (Megaptera novaeangliae) calf riding on mother's head. Near Nomuka Island, Ha'apai Islands, Tonga.

Edwin Giesbers / Red deer (Cervus elaphus) silhouetted stag bellowing during rut in grassland, doe in background. Richmond Park, London, England, UK. October

Andy Rouse / Spotted Hyena (Crocuta crocuta) fighting over hippo carcass in water, Masai Mara, Kenya

Tony Heald / Elephant (Loxodonta africana) reaching up on back legs to feed on tree, front legs on termite mound. Mana Pools National Park, Zimbabwe.

Guy Edwardes / Dalmatian pelican (Pelecanus crispus), portrait. Lake Kerkini, Sintik, Greece. January.

Alex Mustard / Oceanic whitetip shark (Carcharhinus longimanus) swimming close to surface. Brothers Islands, Egypt.

Andy Rouse / Masai Giraffe (Giraffa camelopardalis) silhouetted at sunset, Masai Mara, Kenya

Guy Edwardes / Marbled white (Melanagria galathea), two butterflies on Pyramidal orchid (Anacamptis pyramidalis). Badbury Rings, Wimborne Minster, Dorset, England, UK. June.

Andy Rouse / African Lion (Panthera leo) male at sunrise, Masai Mara, Kenya

Alex Mustard / Jellyfish (Catostylus sp.) swims beneath a slick of plastic debris. Indian Ocean off Sri Lanka. Highly commended in the Environmental Report section, Montphoto competition 2019

Danny Green / Bald eagle (Haliaeetus leucocephalus) in flight, Alaska, USA, February