Nick Garbutt,Nick Garbutt / Tourist watching masses of butterflies drinking from temporary pools. Various species including Large Orange Sulphur (Phoebis agarithe), Cloudless Sulphur (Phoebis sennae), White-angled Sulphur (Anteo...

Anup Shah / Sumatran orangutan (Pongo abelii) female 'Jaki' aged 16 years with baby daughter 'Jodi' aged 2-3 years - portrait. Gunung Leuser National Park, Sumatra, Indonesia. Rehabilitated and released (or desce...

Angelo Gandolfi / Toco Toucan (Ramphastos toco) beak open with tongue visible while feeding on mango, Brazil

Alex Mustard,Alex Mustard / Pair of divers floating in Silfra Canyon, a deep fault filled with fresh water in the rift valley between the Eurasian and American tectonic plates) at Thingvellir National Park, Iceland. May 2011. I...

Inaki Relanzon / Long exposure of Godafoss waterfall, Iceland.

Alex Hyde / Malaysian Orchid Mantis (Hymenopus coronatus) white colour morph camouflaged on an orchid. Captive. Originating from Malaysia.

Erlend Haarberg,Erlend Haarberg / Aerial view down river to the east of the Hofsjakull glacier and west of the Sprengisandur upland desert. Innumerable rivulets flow from beneath the glacier in this area of tundra which abounds in pon...

Andy Rouse / African Lion (Panthera leo) male emerging from bush, Masai Mara, Kenya

Sandesh Kadur / A vine growing on Eucalyptus bark, Tompotika Peninsula, Sulawesi, Indonesia.

Theo Bosboom / Beech trees (Fagus sylvatica) in autumn, seen from under bubbling brook, the Netherlands, November. Runner up in Creative Visions category of the Wildlife Photographer of the Year 2013 competition

Roger Powell / Satellite male Ruff (Philomachus pugnax) in full display at the lek. Varanger, Finmark, Norway, May.

Alex Mustard / Blue shark (Prionace glauca) swimming near the surface of the English Channel. Penzance, Cornwall, England, British Isles. North East Atlantic Ocean, August. Highly commended in the Portraits category...

Chris & Monique Fallows / Great white shark (Carcharodon carcharias) breaching whilst attacking seal decoy, Seal Island, False Bay South Africa.

Milan Radisics / Falconer with Peregrine falcon (Falco peregrinus) to keep other bird away from airport, Budapest Airport, Hungary, March 2009

Jack Dykinga / Rain spattered inter-locking palm leaves create a design of pattern and colour in the tropical rainforest. Osa Peninsula, Costa Rica

Theo Bosboom / Reindeer (Rangifer tarandus) male walking across beach in winter, near Hofn, Iceland, February

Andy Rouse / Maasai Giraffe (Giraffa camelopardalis tippelskirchi) silhouetted by beams of light from setting sun, Maasai Mara, Kenya, Africa. Winner of the African section of the Natures Best Windland Smith Rice...

Diane McAllister,Diane McAllister / Japanese Macaque (Macaca fuscata) male and female huddle together during a light snowfall in Jigokudani Japan, January

Erlend Haarberg / Mountain hare (Lepus timidus) portrait. Vauldalen, Sor-Trondelag, Norway. Highly commended in the GDT 2013 Competition.

Pal Hermansen / Red squirrel (Sciurus vulgaris) sitting on an old car, Bastnas, Sweden, January. Winner of the Fritz Polking Prize at the GDT competition 2013 and winner of the Portfolio category in the Melvita Natur...

Anup Shah / Eastern white-bearded wildebeest (Connochaetes taurinus) herd running. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Ernie Janes / Bengal Tiger (Panthera tigris tigris) captive, with leaves digitally added.

Steven Kazlowski / Polar bear (Ursus maritimus) curious young male on the newly frozen pack ice, Beaufort Sea, off the 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska

Juan Carlos Munoz / Black howler monkeys (Alouatta caraya) male and two females calling from tree, captive, Apenheul Park, Netherlands.

Claudio Contreras / California condor (Gymnogyps californianus), IUCN Critically Endangered, captive.

Tui De Roy / People watching the Fissure eruption of Chico volcano into 9km diameter caldera. Sierra Negra volcano, Isabela Island, Galapagos, Ecuador. October 2005.

Konrad Wothe / Gliding seed of Climbing Gourd (Alsomitra macrocarpa) in tropical rainforest, West Papua, New Guinea, Indonesia, digital composite

Andy Rouse / Giant Panda (Ailuropoda melanoleuca) cub climbing tree. Chengdu, China. Captive.

Mark Bowler,Mark Bowler / Red eyed tree frog (Agalychnis callidryas) walking, digital composite Captive

Suzi Eszterhas / White-faced Capuchin (Cebus capucinus imitator) mother and baby. Osa Peninsula, Costa Rica

Ben Hall / Mute swan (Cygnus olor) stretching its wings backlit at dawn, Poynton, Cheshire, UK, December. Highly honoured in the Bird Category of the Windland Awards of Natures Best Photography Competition 2016...

Florian Möllers / White stork (Ciconia ciconia) adult portrait, captive, Vogelpark Marlow, Germany, May.

Andy Rouse,Andy Rouse / Ring tailed Lemurs (Lemur catta) silhouetted, Madagascar.

Staffan Widstrand / Wild Wonders of China / Low's cymbidium (Cymbidium lowianum) endemic tree orchid, Gaoligongshan NP, Yunnan province, China

Ben Cranke,Ben Cranke / Japanese Macaque (Macaca fuscatata) juvenile, reaching out to touch the camera, Jigokudani, Nagano Prefecture, Honshu, Japan

Juan Carlos Munoz,Juan Carlos Munoz / Leopard (Panthera pardus) portrait, Samburu National Reserve, Kenya

Dave Watts,Dave Watts / Forester Kangaroo family {Macropus giganteus tasmaniensis} a subspecies of Eastern Grey Kangaroo, Tasmania, Australia.

Andrew Parkinson / Gannet (Morus bassanus) breeding pair during part of their elaborate courtship ritual. Saltee Islands, Republic of Ireland, May.

Yukihiro Fukuda,Yukihiro Fukuda / Japanese fireflies (Luciola cruciata) in flight at night, endemic species, Yaku-shima UNESCO World Heritage Site, Kagoshima, Japan, May

Anup Shah,Anup Shah / Lioness (Panthera leo) resting on a fallen tree with a courting male. Masai Mara National Reserve, Kenya, July

Juan Carlos Munoz,Juan Carlos Munoz / Starfish Beach, with many starfish in the shallow sea (Asteroidea) Colon Island, Bocas del Toro Archipelago, Bocas del Toro Province, Panama,

Juan Carlos Munoz / Gemsbok (Oryx gazella) silhouetted at dawn, Kalahari Desert, Botswana.

Juan Carlos Munoz / Two Great egrets (Ardea alba) wading, silhouetted at dawn, Keoladeo National Park, Bharatpur, Rajasthan, India.

Juergen Freund / Bioluminescent fungi (Omphalotus nidiformis / Pleurotus nidiformis) glowing on tree trunk in rainforest at night, Atherton Tablelands, Queensland, Australia,

Andy Rouse,Andy Rouse / African Elephant (Loxodonta africana) breeding herd led by matriarch. Maasai Mara, Kenya, Africa, August.

Mark Bowler / Isula / Bullet ant (Paraponera clavata) close-up showing mandibles, Peru

Enrique Lopez-Tapia,Enrique Lopez-Tapia / Broad Peak base camp (4,960 m) at night, with K2 (8,611m) and the Godwin-Austen glacier in the background lit by moonlight, Central Karakoram National Park, Pakistan, June 2007. Winner of Photographer...

Andy Rouse,Andy Rouse / European Otter (Lutra lutra) running. Controlled conditions. Sussex, UK, October.

Mark MacEwen / Lioness (Panthera leo) feeding on carcass, Ngorongoro Crater, Tanzania, Africa.

Benjamin Barthelemy,Benjamin Barthelemy / Ibex (Capra Ibex) fighting, Aiguilles Rouges National Nature Reserve, Haute-Savoie, France, June

Benjamin Barthelemy,Benjamin Barthelemy / European White water lily, (Nymphaea alba) in flowers, Vallee de l'Eure, Eure-et-Loir, France, July.

Sven Zacek,Sven Zacek / Roe deer (Capreolus capreolus) bucks on snowy field in SOuthern Estonia, March.

Alex Mustard / A long exposure renders Bar jacks (Carangoides ruber) as ghostly trails as they swim around a sponge, on a coral reef, at night. East End, Grand Cayman, Cayman Islands. Overall winner of GDT competiti...

Ernie Janes,Ernie Janes / Newly hatched ducklings with eggs.

Alex Mustard,Alex Mustard / A signal blenny (Emblemaria hypacanthus) displays its showy dorsal fin as it extends out of its tube home. La Paz, Mexico. Sea Of Cortez, East Pacific Ocean.

Theo Bosboom,Theo Bosboom / White-legged damselfly (Platycnemis pennipis), Meinerswijk, The Netherlands, June 2005. Highly commended, Animal portraits category, Wildlife Photographer of the Year Awards 2006.

Radomir Jakubowski / Marbled white butterfly (Melanargia galathea) on Horsetail (Equisetum), Herbitzheim near Saarbruecken, Saarland, Germany. Finalist in the Macro category and winner of the audience award in the Montpho...

Adrian Davies / Fossilised fly in Baltic amber. Whole specimen approx 15mm long. Fly approx 5mm.

Sue Daly,Sue Daly / Jewel Anemones (Corynactis haddoni) colonising the wreck of HMNZS Canterbury, Bay of Islands, New Zealand, February 2013

Juergen Freund / Forest seedling growing in the Bornean forest. Gunung Gading National Park, Sarawak, Malaysian Borneo