Alex Mustard / Hedgehog seahorse (Hippocampus spinosissimus) female moves over a sandy seabed. Dauin, Negros Island, Philippines. Bohol Sea, tropical west Pacific Ocean.

Alex Mustard / Spaghetti garden eels (Gorgasia maculata) stretching up out of their burrows on a rubble slope, with Mysiid shrimp swimming around their bodies near the seabed. Dumaguete, Dauin, Negros, Philippines....

Alex Mustard / Portrait of Morgan's coral hermit crab (Paguritta morgani) looking out of its burrow in a hard coral colony. These hermit crabs are sedentary, remaining in their burrows, rather than carrying arou...

Alex Mustard / Interaction, possibly predation, between two polychaete worms (Chloeia sp). Dauin, Dauin Marine Protected Area, Dumaguete, Negros, Philippines. Bohol Sea, tropical west Pacific Ocean.

Alex Mustard / Bue ring octopus (Hapalochlaena lunulata) walking across the sea bed. Dauin, Dauin Marine Protected Area, Dumaguete, Negros, Philippines. Bohol Sea, tropical west Pacific Ocean.

Alex Mustard / Nudibranch (Nembrotha milleri) on the seabed. Dauin, Dauin Marine Protected Area, Dumaguete, Negros, Philippines. Bohol Sea, tropical west Pacific Ocean.

Alex Mustard / Blue ring octopus (Hapalochlaena lunulata) walks across the sea bed. Dauin, Dauin Marine Protected Area, Dumaguete, Negros, Philippines. Bohol Sea, tropical west Pacific Ocean.

Alex Mustard / Wonderpus octopus (Wunderpus photogenicus) reared up on its long arms for a better view. Dauin, Dauin Marine Protected Area, Dumaguete, Negros, Philippines. Bohol Sea, tropical west Pacific Ocean.

Alex Mustard / Portrait of a Tropical bottletail squid (Sepiadarium kochi) partially buried in sand and illuminated in a beam of light, at night. Dauin, Dauin Marine Protected Area, Dumaguete, Negros, Philippines. B...

Alex Mustard / Large blue ring octopus (Hapalochlaena lunulata) swimming just above the sea bed. Dauin, Dauin Marine Protected Area, Dumaguete, Negros, Philippines. Bohol Sea, tropical west Pacific Ocean.

Alex Mustard / Warty frogfish (Antennartius maculatus) female raising her elaborate lure to attract prey. Dauin, Dauin Marine Protected Area, Dumaguete, Negros, Philippines. Bohol Sea, tropical west Pacific Ocean.

Alex Mustard / Box crab (Calappa sp.) female emerging from the sand and shakes her body to release her tiny young. Dauin, Dauin Marine Protected Area, Dumaguete, Negros, Philippines. Bohol Sea, tropical west Pacific...

Alex Mustard / Pair of pygmy lemon gobies (Lubricogobius exiguus) make their home in an old heart urchin (Maretia sp.) test. Dauin, Dauin Marine Protected Area, Dumaguete, Negros, Philippines. Bohol Sea, tropical we...

Alex Mustard / Pink anemonefish (Amphiprion perideraion) tends a clutch of new eggs (revealed by their red colour) beneath its host anemone (magnificent sea anemone: Heteractis magnifica). Dauin, Dumaguete, Negros,...

Alex Mustard / Pink anemonefish (Amphiprion perideraion) fans its clutch of eggs with its pectoral fin (these eggs are aged 3-4 days), beneath its Magnificent sea anemone (Heteractis magnifica). Dauin, Dumaguete, Ne...

Alex Mustard / Threadfin anthias (Pseudanthias huchti) males locking jaws as they fight over territory on a coral reef. Dauin, Dauin Marine Protected Area, Dumaguete, Negros, Philippines. Bohol Sea, tropical west Pa...

Alex Mustard / Ringtailed cardinalfish (Apogon aureus) male mouthbrooding eggs. The colour of these eggs shows that they are close to hatching. Dauin, Dumaguete, Negros, Philippines. Bohol Sea, tropical west Pacific...

Alex Mustard / Freshwater terrapin (Trachemys scripta elegans) in a garden pond. Dauin, Dumaguete, Negros, Philippines.

Alex Mustard / Hearth heart urchin (Maretia planulata) which usually lives buried in the sand, crawls along the seabed. Dauin, Dumaguete, Negros, Philippines. Bohol Sea, tropical west Pacific Ocean.

Alex Mustard / Seapens (Virgularia sp.) growing in shallow water on seabed of muddy sediment. Dauin, Negros Island, Philippines. Bohol Sea, tropical west Pacific Ocean.

Alex Mustard / Seapens (Virgularia sp.) emerging from the sediment of a muddy slope. Dauin, Negros Island, Philippines. Bohol Sea, tropical west Pacific Ocean.

Alex Mustard / False clown anemonefish (Amphiprion oceallaris) male tending his eggs, cleaning them with his mouth and aerating them by beating his pectoral fins. Dauin, Dumaguete, Negros, Philippines. Bohol Sea, T...

Alex Mustard / False clown anemonefish (Amphiprion oceallaris) eggs that are close to hatching. Dauin, Dumaguete, Negros, Philippines. Bohol Sea, Tropical West Pacific Ocean.

Alex Mustard / Algae octopus (Abdopus aculeatus) moves over a sandy seabed, Dauin, Negros Island, Philippines. Bohol Sea, tropical west Pacific Ocean.

Alex Mustard / Mozambique scorpionfish (Parascorpaena mossambica) two males disputing territory on the seabed, yawning is a threat display, Dauin, Dumaguete, Negros, Philippines. Bohol Sea, tropical west Pacific Oce...

Alex Mustard / Hairy / Straited frogfish (Antennarius striatus) pair in their typical habitat, male on left, female on right panting with a belly fully of eggs. Dauin, Negros Island, Philippines. Bohol Sea, tropical...

Alex Mustard / Commensal emperor shrimp (Periclimenes imperator / Zenopontonia rex) scuttles across the skin of its host sea cucumber, Dauin, Dumaguete, Negros, Philippines. Bohol Sea

Alex Mustard / Ornate ghost pipefish (Solenostomus paradoxus) the larger female is above, her pelvic fins are cupping a clutch of eggs, male below, Dauin, Negros Island, Philippines. Bohol Sea, Pacific Ocean.

Alex Mustard / Orange painted frogfish (Antennarius pictus) tries to attract prey using its esca (lure) Dauin, Dumaguete, Negros, Philippines. Bohol Sea, Pacific Ocean.

Alex Mustard / Flamboyant cuttlefish (Metasepia pfefferi) on the sand, Dauin, Negros Island, Philippines. Bohol Sea, Pacific Ocean.

Alex Mustard / Orange hairy / Straited frogfish (Antennarius striatus) yawns while illuminated by a snooted strobe, creating a spotlight. Frogfish yawn as an aggressive behaviour. Dauin, Negros Island, Philippines....

Alex Mustard / Flamboyant cuttlefish (Metasepia pfefferi) crawling over sand, Dauin, Dumaguete, Negros, Philippines. Bohol Sea, Pacific Ocean.

Alex Mustard / Harlequin swimming crab (Lissocarcinus laevis) sheltering beneath the tentacles of a tube anemone (Cerianthus sp.). Dauin, Dumaguete, Negros, Philippines. Bohol Sea, Tropical West Pacific Ocean.

Alex Mustard / Long exposure of a juvenile Harlequin sweetlips (Plectorhinchus chaetodonoides), which dance as a defence from predators. The dance may mimic the movements of poisonous flatworms or may simply confuse...

Alex Mustard / Clark's anemonefish (Amphiprion clarkii) at host anemone. Dauin, Dumaguete, Negros, Philippines. Bohol Sea, Tropical West Pacific Ocean. Parent of the eggs in image 1527884

Alex Mustard / Clark's anemonefish (Amphiprion clarkii) eggs at host anemone. Dauin, Dumaguete, Negros, Philippines. Bohol Sea, Tropical West Pacific Ocean. Parent of the eggs in image 1527885

Alex Mustard / Clark's anemonefish (Amphiprion clarkii) eggs in late stage development, laid between folds in an anemone. Dauin, Dumaguete, Negros, Philippines. Bohol Sea, Tropical West Pacific Ocean.

Alex Mustard / False clown anemonefish (Amphiprion ocellaris) looks out from an anemone. Dauin, Dumaguete, Negros, Philippines. Bohol Sea, Tropical West Pacific Ocean.

Alex Mustard / Devil scorpionfish (Inimicus didactylus) moves across sandy seabed. Dauin, Dumaguete, Negros, Philippines. Bohol Sea, Tropical West Pacific Ocean.

Alex Mustard / Group of Saddleback anemonefish (Amphiprion polymnus) guard clutch of eggs, laid on stone near the base of anemone. Dauin, Dumaguete, Negros, Philippines. Bohol Sea, Tropical West Pacific Ocean.

Alex Mustard / Pair of male Longtail dragonets (Callionymus neptunius) aggressively displaying to each other, Dauin, Dumaguete, Negros, Philippines. Bohol Sea, Tropical West Pacific Ocean.

Alex Mustard / Blue ringed octopus (Hapalochlaena lunulata) displaying as it moves across the seabed. Dauin, Dumaguete, Negros, Philippines. Bohol Sea, Tropical West Pacific Ocean.

Alex Mustard / Blue ringed octopus (Hapalochlaena lunulata) displays as it moves across the seabed. Dauin, Dumaguete, Negros, Philippines. Bohol Sea, Tropical West Pacific Ocean.

Alex Mustard / Dragon shrimp (Micropandalus hardingi) resting on coral, portrait, Dauin Marine Protected Area, Negros, Philippines, Bohol Sea, Pacific Ocean.

Alex Mustard / Orange painted frogfish (Antennarius pictus) hiding on the seabed, next to photographers thumb for scale, Dauin Marine Protected Area, Philippines, Bohol Sea, Pacific Ocean.

Alex Mustard / Ornate ghost pipefish (Solenostomus paradoxus), juvenile, hides within a crinoid, Dauin Marine Protected Area, Dumaguete, Negros, Philippines. Wildlife Photographer of the Year 2021 competititon cat...

Alex Mustard / RF - Pair of Pink anemonefish (Amphiprion perideraion) look out from within their host Magnificent sea anemone (Heteractis magnifica). Dauin, Dauin Marine Protected Area, Dumaguete, Negros, Philippine...

Alex Mustard / Two photos of the same Algae octopus (Abdopus aculeatus) showing how quickly is can change its shape, colour and texture. Dauin, Dauin Marine Protected Area, Dumaguete, Negros, Philippines. Bohol Sea,...

Alex Mustard / Portrait of a large female Emperor shrimp (Zenopontonia rex/ Periclimenes imperator) on a sea cucumber (Bohadschia sp.). Dauin, Dauin Marine Protected Area, Dumaguete, Negros, Philippines. Bohol Sea,...

Alex Mustard / Portrait of a Nudibranch (Nembrotha milleri) on a coral reef. Dauin, Dauin Marine Protected Area, Dumaguete, Negros, Philippines. Bohol Sea, tropical west Pacific Ocean.

Alex Mustard / Pink anemonefish (Amphiprion perideraion) cares for its eggs beneath a magnificent sea anemone (Heteractis magnifica) on a coral reef. Dauin, Dauin Marine Protected Area, Dumaguete, Negros, Philippine...

Alex Mustard / Wide angle macro photo of an algae octopus (Abdopus aculeatus) as it reaches out and investigates the camera. Dauin, Dauin Marine Protected Area, Dumaguete, Negros, Philippines. Bohol Sea, tropical we...

Alex Mustard / Portrait of a Black saddle snake eel (Ophichthus cephalozona) emerging from the sand at night. Dauin, Dauin Marine Protected Area, Dumaguete, Negros, Philippines. Bohol Sea, tropical west Pacific Ocea...

Alex Mustard / Detail of rhinophores and toxic lack markings on a Funeral nudibranch (Jorunna funebris). Dauin, Dauin Marine Protected Area, Dumaguete, Negros, Philippines. Bohol Sea, tropical west Pacific Ocean.

Alex Mustard / Common ghostgoby (Pleurosicya mossambica) hiding on sponge. Dauin Marine Protected Area, Dumaguete, Negros, Philippines. Bohol Sea, tropical west Pacific Ocean.

Alex Mustard / Yellow phantom velvetfish (Paraploactis kagoshimensis) rests on the sea bed. Dauin, Dauin Marine Protected Area, Dumaguete, Negros, Philippines. Bohol Sea, tropical west Pacific Ocean.

Alex Mustard / Tiny skinspot dwarfgoby (Trimma halonevum) on seabed. Dauin, Dauin Marine Protected Area, Dumaguete, Negros, Philippines. Bohol Sea, tropical west Pacific Ocean.

Alex Mustard / Long exposure of a school of Black-axil chromis (Chromis atripectoralis) swimming in formation against a current. Dauin, Dauin Marine Protected Area, Dumaguete, Negros, Philippines. Bohol Sea, tropica...

Alex Mustard / Portrait of a Clark's anemonefish (Amphiprion clarkii) in a sea anemone on a coral reef. Dauin, Dauin Marine Protected Area, Dumaguete, Negros, Philippines. Bohol Sea, tropical west Pacific Ocean.

Alex Mustard / Shallow depth of field photo of an adult saddleback anemonefish (Amphiprion polymnus) fanning eggs, laid on a stone near the base of the anemone. Dauin, Dumaguete, Negros, Philippines. Bohol Sea, Trop...

Alex Mustard / Pair of Reticulated dascyllus (Dascyllus retriculatus) spawning eggs on a coral reef. Dauin, Dauin Marine Protected Area, Dumaguete, Negros, Philippines. Bohol Sea, tropical west Pacific Ocean.

Alex Mustard / Flamboyant cuttlefish (Metasepia pfefferi) female extending her arms to catch prey (usually small crustaceans). Dauin, Dauin Marine Protected Area, Dumaguete, Negros, Philippines. Bohol Sea, tropical...

Alex Mustard / Saddleback anemonefish (Amphiprion polymnus) barks a warning as it guards a clutch of developing eggs (beneath it) laid on a small stone, below a fold in its home, a Haddon's sea anemone (Stichoda...

Alex Mustard / A colourful flamboyant cuttlefish (Metasepia pfefferi) with a captured shrimp in its tentacles. Dauin, Dauin Marine Protected Area, Dumaguete, Negros, Philippines. Bohol Sea, tropical west Pacific Oce...

Alex Mustard / Portrait of a Giant frogfish (Antennarius commersoni) waiting in ambush hidden amongst sponges growing on a coral reef. Dauin Marine Protected Area, Dumaguete, Negros, Philippines. Bohol Sea, tropical...

Alex Mustard / Dragon shrimp (Micropandalus hardingi) on a coral reef. Dauin Marine Protected Area, Dumaguete, Negros, Philippines. Bohol Sea, tropical west Pacific Ocean.

Alex Mustard / Tigertail seahorse (Hippocampus comes) female, Dauin Marine Protected Area, Dumaguete, Negros, Philippines. Bohol Sea, tropical west Pacific Ocean.

Alex Mustard / Orange short-tailed pipefish (Trachyrhamphus bicoarctatus) on ropes on the seabed. Dauin Marine Protected Area, Dumaguete, Negros, Philippines. Bohol Sea, tropical west Pacific Ocean.