Floris van Breugel / Natural iridescent oil film from decaying leaves at the surface of a pool, with a cottonwood leaf floating. Escalante National Monument, Utah. November.

Klaus Echle / After being trapped in an empty shed, Badger (Meles meles) passes wall decorated with graffiti on its way back to the forest. Baden-Wuerttemburg, Germany, May 2010. Winner of the Man and Nature catego...

Juergen Freund / Very old human skulls on a ledge in the karst limestone walls of a Raja Ampat island. Raja Ampat, West Papua, Indonesia, February 2010

Laurent Geslin / Common bee fly (Bombylius major) at rest with graffiti in the background, Paris, France.

Michael Hutchinson / Groundsel {Senecio sp} growing out of brick wall covered in colourful graffiti, Bristol, UK

Guy Edwardes / Close up of decaying Sycamore leaf (Acer pseudoplatanus) Devon, England, November 2008

Nick Upton / Yellow dung fly (Scathophaga stercoraria) killed by pathogenic fungus of flies (Entomophthora muscae) clinging to a thistle leaf with wings spread, Wiltshire pastureland, UK, June.

Alex Hyde / Fruiting body of fungi (Crepidotus applanatus / variabilis) growing on leaf litter in beech woodland. Peak District National Park, Derbyshire, UK, October.

Fabrice Cahez / Dead trunks overgrown with moss. Vosges mountain, France, October.

Loic Poidevin / Moss-covered dead tree, near rapids 'du pas de l'ours'. French Pyrenees, September 2012.

Adrian Davies / Peniophora quercina (Peniophora quercina), a wood decay fungus, growing on tree bark, Surrey, England, UK, October.

Floris van Breugel / Natural iridescent plant oils from decaying leaves at the surface of a pool, Escalante National Monument, Utah. November

Will Watson / Natural oil film on surface of woodland pond, caused by the anoxic decay of organic matter, Essex, England, UK, March

Philippe Clement / Fallen tree trunk left to rot on forest floor as dead wood habitat for invertebrates, Pyrenees, France

Sandra Bartocha / European Beech (Fagus sylvatica) two decaying leaves against dead wood, Germany, April

Peter Cairns / 2020VISION / Decaying sycamore leaf. Scotland, October.

Alex Hyde / Dung Cup (Peziza vesiculosa) fungal fruiting bodies growing on horse manure. Derbyshire, UK, April.

David Shale / Decomposing whale bone and epifauna. Collected from coral sea mount near Dragon vent field on SW Indian Ridge, Indian Ocean.

David Shale / Decomposing whale bone and epifauna. Collected from coral sea mount near Dragon vent field on SW Indian Ridge, Indian Ocean.

David Shale / Bivalve molluscs on whale bone. Collected from coral sea mount near Dragon vent field on SW Indian Ridge, Indian Ocean.

David Shale / Epifauna on whale bone. Collected from coral sea mount near Dragon vent field on SW Indian Ridge, Indian Ocean.

Danny Green / Slow Worm (Anguis fragilis) basking on rusty metal. Leicestershire, UK, March.

Danny Green / Slow Worm (Anguis fragilis) basking on rusty metal. Leicestershire, UK, March.

Guy Edwardes / 2020VISION / Toadstools growing from falled trunk. New Forest National Park, Hampshire, UK, October.

Adrian Davies / Ochre cushion fungus (Hypocrea pulvinata) growing on old fruit body of Birch Polypore (Piptoporus betulinus) England, December.

Brent Stephenson / Kelp Gull (Larus dominicanus) chick amongst old whaling equipment. Deception Island, South Shetland Islands, Antarctica, January.

Alex Hyde / Sulphur Polypore / Chicken of the Woods (Laetiporous sulphureus) growing on tree trunk. Nordtirol, Tirol, Austrian Alps, Austria, 2300 metres, July.

Alex Hyde / Sulphur Polypore / Chicken of the Woods (Laetiporous sulphureus) growing on tree trunk. Nordtirol, Tirol, Austrian Alps, Austria, 2300 metres, July.

Anup Shah / Carcass of Eastern White bearded wildebeest(Connochates taurinus albojubatus) covered in maggots, Masai Mara National Reserve, Kenya

Anup Shah / Carcass of Eastern White bearded wildebeest(Connochates taurinus albojubatus) covered in maggots, Masai Mara National Reserve, Kenya

Gary K. Smith / Fungi, Dryads Saddle (Polyporus squamosus) fresh fruiting bodies on fallen tree stump. Norfolk, UK, May.

Peter Cairns / Gnarled Scots Pine (Pinus sylvestris) tree framing western Grampians crowned with heavy clouds. Cairngorms National Park, Scotland, May.

Nick Upton / Horse's hoof / Tinder fungus (Fomes fomentarius) growing on European beech (Fagus sylvatica) tree trunk, Plitvice Lakes National Park, Croatia, July.

Nick Garbutt / Veins and filaments of a leaf after decomposition. Lowland rainforest, Masoala National Park, Madagascar.

Tim Laman / Strangler Fig (Ficus sp.) tree with decomposing host tree visible in the center. Lowland rainforest in Borneo. Gunung Palung National Park, Indonesia.

Nick Upton / Close up of Greenbottle / Blowfly (Lucilia sp.) inspecting a dead mole, before laying eggs on it. Wiltshire garden, UK, June.

Nick Upton / Greenbottle / Blowfly (Lucilia sp.) inspecting a dead mole, before laying eggs on it. Wiltshire garden, UK, June.

Nick Upton / Greenbottle / Blowfly (Lucilia sp.) inspecting a dead mole, before laying eggs on it. Wiltshire garden, UK, June.

Nick Upton / Greenbottle / Blowfly (Lucilia sp.) inspecting a dead mole, before laying eggs on it. Wiltshire garden, UK, June.

Dietmar Nill / Cross section of European beech trunk (Fagus sylvaticus) showing fungal decay, Germany

Gerrit Vyn / Downy Woodpecker (Picoides pubescens) female, foraging on a decaying tree. February, Tompkins County, New York

Guy Edwardes / Close up of decaying Sycamore leaf (Acer pseudoplatanus) Devon, England, November 2008

Guy Edwardes / Close up of decaying Sycamore leaf (Acer pseudoplatanus) Devon, England, November 2008

Paul Hobson / Oak leaves {Quercus robur} in leaf litter in autumn, Derbyshire, UK

Nick Garbutt / Fungi (unknown sp) growing on fallen log on forest floor, Danum Valley, Sabah, Borneo, Malaysia

Jerry Monkman / Fallen logs decay into the mossy forest floor in old spruce forest, Isle Au Haut, Acadia National Park, Maine, USA, The Duck Harbor Trail.

Niall Benvie / Shaggy inkcap fungus / Lawyer's wig {Coprinus comatus} deliquescing, Norway

Jane Burton / Common frog {Rana temporaria} erythristic tadpoles with normal tadpoles around a skeletonised oak leaf.

WILLIAM OSBORN / Fallen Scots Pine tree with holes from wood boring insects Pinus sylvestris}