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Cicada turret, or chimney, formed by underground larva. Sabah, Borneo.
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Adrian Davies / Cicada turret, or chimney, formed by underground larva. Sabah, Borneo.
Baboon spoor. Moremi reserve, Botswana, Southern Africa
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Richard Du Toit / Baboon spoor. Moremi reserve, Botswana, Southern Africa
Marsican / Apennine brown bear (Ursus arctos marsicanus) claw marks on Beech (Fagus sylvatica) tree bark in old growth beech forest. Abruzzo, Lazio and Molise National Park / Parco Nazionale d'Abr...
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Bruno D'Amicis / Marsican / Apennine brown bear (Ursus arctos marsicanus) claw marks on Beech (Fagus sylvatica) tree bark in old growth beech forest. Abruzzo, Lazio and Molise National Park / Parco Nazionale d'Abr...
Hazel nut (Corylus avellana) cases on forest floor, broken by grey squirrel. Dorset, UK August.
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Colin Varndell / Hazel nut (Corylus avellana) cases on forest floor, broken by grey squirrel. Dorset, UK August.
Polar Bear footprints in melting permafrost on Northern Svalbard. July 2013
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Ashley Cooper / Polar Bear footprints in melting permafrost on Northern Svalbard. July 2013
Goat willow (Salix caprea) felled by a Eurasian beaver (Castor fiber) with teeth marks The riverside tree will resprout with new shoots supplying a source of food for the beavers for years to come. De...
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Matthew Maran / Goat willow (Salix caprea) felled by a Eurasian beaver (Castor fiber) with teeth marks The riverside tree will resprout with new shoots supplying a source of food for the beavers for years to come. De...
Goat willow (Salix caprea) felled by a Eurasian beaver (Castor fiber). The riverside tree will resprout with new shoots supplying a source of food for the beavers for years to come. Devon, England, UK...
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Matthew Maran / Goat willow (Salix caprea) felled by a Eurasian beaver (Castor fiber). The riverside tree will resprout with new shoots supplying a source of food for the beavers for years to come. Devon, England, UK...
Siberian tiger (Panthera tigris tigris) scat, Sikhote-Alin Mountain Range, Sikhote-Alin Nature Reserve, Primorye Krai, Russian Far East. May.
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Igor Shpilenok / Siberian tiger (Panthera tigris tigris) scat, Sikhote-Alin Mountain Range, Sikhote-Alin Nature Reserve, Primorye Krai, Russian Far East. May.
Hemlock water dropwort stem (Oenanthe crocata) cut by a Water vole (Arvicola amphibius) at a tell-tale 45 degree angle, found during a survey for signs of Water vole activity.  This plant is highly to...
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Nick Upton / Hemlock water dropwort stem (Oenanthe crocata) cut by a Water vole (Arvicola amphibius) at a tell-tale 45 degree angle, found during a survey for signs of Water vole activity. This plant is highly to...
Footprints of Water voles (Arvicola amphibius), or possibly Brown rats (Rattus norvegicus) pointed out on a muddy river bank, found during a survey for signs of Water voles, near Bude, Cornwall, UK, A...
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Nick Upton / Footprints of Water voles (Arvicola amphibius), or possibly Brown rats (Rattus norvegicus) pointed out on a muddy river bank, found during a survey for signs of Water voles, near Bude, Cornwall, UK, A...
Erect-crested penguins (Eudyptes sclateri) feathers left behind after seasonal moulting. Antipodes Island, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered specie...
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Tui De Roy / Erect-crested penguins (Eudyptes sclateri) feathers left behind after seasonal moulting. Antipodes Island, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered specie...
Erect-crested penguins (Eudyptes sclateri) feathers left behind after seasonal moulting. Antipodes Island, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered specie...
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Tui De Roy / Erect-crested penguins (Eudyptes sclateri) feathers left behind after seasonal moulting. Antipodes Island, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered specie...
Rhino rubbing rock, iMfolozi National Park, South Africa
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Staffan Widstrand / Rhino rubbing rock, iMfolozi National Park, South Africa
Rhino rubbing post  rhinos, both black and white rhinos. They scratch themselves on this tree, since decades back. iMfolozi National Park, South Africa
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Staffan Widstrand / Rhino rubbing post rhinos, both black and white rhinos. They scratch themselves on this tree, since decades back. iMfolozi National Park, South Africa
Common birch pygmy moth (Stigmella betulicola) larva mine in Silver Birch leaf.  Surrey, England, UK.
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Adrian Davies / Common birch pygmy moth (Stigmella betulicola) larva mine in Silver Birch leaf. Surrey, England, UK.
Common birch pygmy moth (Stigmella betulicola) mine in Silver Birch leaf.  Surrey, England, UK.
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Adrian Davies / Common birch pygmy moth (Stigmella betulicola) mine in Silver Birch leaf. Surrey, England, UK.
Scarlet caterpillar club (Cordyceps militaris) a parasitic fungus on the buried larvae of butterflies and moths. Sussex, England, UK, November.
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Adrian Davies / Scarlet caterpillar club (Cordyceps militaris) a parasitic fungus on the buried larvae of butterflies and moths. Sussex, England, UK, November.
European bison (Bison bonasus) damage on trees, causde by rubbing against the bark, Zuid-Kennemerland National Park,  the Netherlands. January. Reintroduced species.
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Edwin Giesbers / European bison (Bison bonasus) damage on trees, causde by rubbing against the bark, Zuid-Kennemerland National Park, the Netherlands. January. Reintroduced species.
Spotted hyaena (Crocuta crocuta) footprints, Senegal.
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Roland Seitre / Spotted hyaena (Crocuta crocuta) footprints, Senegal.
Spotted hyaena (Crocuta crocuta) faeces, Senegal.
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Roland Seitre / Spotted hyaena (Crocuta crocuta) faeces, Senegal.
Downy birch tree (Betula pubescens) heavily gnawed by Eurasian beavers (Castor fiber) and close to being felled, in  large enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, May.
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Nick Upton / Downy birch tree (Betula pubescens) heavily gnawed by Eurasian beavers (Castor fiber) and close to being felled, in large enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, May.
Rear, webbed footprint of Eurasian beaver (Castor fiber) in mud within enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, May.
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Nick Upton / Rear, webbed footprint of Eurasian beaver (Castor fiber) in mud within enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, May.
Toothmarks left by Eurasian beaver (Castor fiber) that has stripped bark from a cut tree branch within a large woodland enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, April 2015.
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Nick Upton / Toothmarks left by Eurasian beaver (Castor fiber) that has stripped bark from a cut tree branch within a large woodland enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, April 2015.
Toothmarks left by Eurasian beaver (Castor fiber) that has stripped bark from a cut tree branch within a large woodland enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, April 2015.
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Nick Upton / Toothmarks left by Eurasian beaver (Castor fiber) that has stripped bark from a cut tree branch within a large woodland enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, April 2015.
Toothmarks of Eurasian beavers (Castor fiber) on Willow tree (Salix sp.) trunk within a large woodland enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, April 2015.
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Nick Upton / Toothmarks of Eurasian beavers (Castor fiber) on Willow tree (Salix sp.) trunk within a large woodland enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, April 2015.
Tree felled by Eurasian beavers (Castor fiber) on the banks of the River Otter, Devon, UK, March.
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Nick Upton / Tree felled by Eurasian beavers (Castor fiber) on the banks of the River Otter, Devon, UK, March.
Willow tree (Salix sp.) heavily gnawed by Eurasian beavers (Castor fiber) within a large woodland enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, April.
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Nick Upton / Willow tree (Salix sp.) heavily gnawed by Eurasian beavers (Castor fiber) within a large woodland enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, April.
Willow tree (Salix sp.) trunk heavily gnawed by Eurasian beavers (Castor fiber) within a large woodland enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, April.
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Nick Upton / Willow tree (Salix sp.) trunk heavily gnawed by Eurasian beavers (Castor fiber) within a large woodland enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, April.
Willow tree (Salix sp.) trunk heavily gnawed by Eurasian beavers (Castor fiber) within a large woodland enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, April.
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Nick Upton / Willow tree (Salix sp.) trunk heavily gnawed by Eurasian beavers (Castor fiber) within a large woodland enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, April.
Willow tree (Salix sp.) heavily gnawed by Eurasian beavers (Castor fiber) within a large woodland enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, April.
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Nick Upton / Willow tree (Salix sp.) heavily gnawed by Eurasian beavers (Castor fiber) within a large woodland enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, April.
Willow tree (Salix sp.) heavily gnawed by Eurasian beavers (Castor fiber) within a large woodland enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, April.
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Nick Upton / Willow tree (Salix sp.) heavily gnawed by Eurasian beavers (Castor fiber) within a large woodland enclosure, Devon Beaver Project, Devon Wildlife Trust, Devon, UK, April.
Willow trees (Salix sp.) gnawed and felled by Eurasian beavers (Castor fiber), Wiltshire, UK, April.
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Nick Upton / Willow trees (Salix sp.) gnawed and felled by Eurasian beavers (Castor fiber), Wiltshire, UK, April.
Young Willow tree (Salix sp.) felled by Eurasian beavers (Castor fiber), Gloucestershire, UK, April.
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Nick Upton / Young Willow tree (Salix sp.) felled by Eurasian beavers (Castor fiber), Gloucestershire, UK, April.
Young Willow trees (Salix sp.) felled and gnawed by Eurasian beavers (Castor fiber), Gloucestershire, UK, April.
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Nick Upton / Young Willow trees (Salix sp.) felled and gnawed by Eurasian beavers (Castor fiber), Gloucestershire, UK, April.
Young Willow tree (Salix sp.) felled by Eurasian beavers (Castor fiber), Gloucestershire, UK, April.
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Nick Upton / Young Willow tree (Salix sp.) felled by Eurasian beavers (Castor fiber), Gloucestershire, UK, April.
Young Willow tree (Salix sp.) felled by Eurasian beavers (Castor fiber), Gloucestershire, UK, April.
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Nick Upton / Young Willow tree (Salix sp.) felled by Eurasian beavers (Castor fiber), Gloucestershire, UK, April.
Brown hare (Lepus europaeus) tracks in the snow, Norfolk, UK, January.
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David Tipling / Brown hare (Lepus europaeus) tracks in the snow, Norfolk, UK, January.
Otter (Lutra lutra) dropping / spraint on rock at edge of lake. Devon, England, July.
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Adrian Davies / Otter (Lutra lutra) dropping / spraint on rock at edge of lake. Devon, England, July.
Close up of plant  stem  cut by Water vole (Arvicola amphibius) at 45 degree angle. Found on lake bank during survey for signs of Water vole activity, near Bude, Cornwall, UK, April 2014.
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Nick Upton / Close up of plant stem cut by Water vole (Arvicola amphibius) at 45 degree angle. Found on lake bank during survey for signs of Water vole activity, near Bude, Cornwall, UK, April 2014.
Rebecca Northey of Derek Gow Consultancy inspecting stems of Stinging nettle (Urtica dioica) cut by Water vole (Arvicola amphibius) and left near its burrow entrance on river bank. Found during survey...
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Nick Upton / Rebecca Northey of Derek Gow Consultancy inspecting stems of Stinging nettle (Urtica dioica) cut by Water vole (Arvicola amphibius) and left near its burrow entrance on river bank. Found during survey...
Dani Siddall of Derek Gow Consultancy looking at stem of Hemlock water dropwort (Oenanthe crocata) cut by Water vole (Arvicola amphibius), and left on river bank. Found during survey for signs of Wate...
01488111
Nick Upton / Dani Siddall of Derek Gow Consultancy looking at stem of Hemlock water dropwort (Oenanthe crocata) cut by Water vole (Arvicola amphibius), and left on river bank. Found during survey for signs of Wate...
Black-backed jackal  (Canis mesomelas) footprints, Dead Vlei, Namib Desert, Namibia, April
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Ann & Steve Toon / Black-backed jackal (Canis mesomelas) footprints, Dead Vlei, Namib Desert, Namibia, April
Black-backed jackal  (Canis mesomelas) footprints, Dead Vlei, Namib Desert, Namibia, April
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Ann & Steve Toon / Black-backed jackal (Canis mesomelas) footprints, Dead Vlei, Namib Desert, Namibia, April
Burrow of Dancing white lady spider (Leucorchestris arenicola), Namib Desert, Namibia, May.
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Ann & Steve Toon / Burrow of Dancing white lady spider (Leucorchestris arenicola), Namib Desert, Namibia, May.
Burrow of Dancing white lady spider (Leucorchestris arenicola), Namib Desert, Namibia, May.
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Ann & Steve Toon / Burrow of Dancing white lady spider (Leucorchestris arenicola), Namib Desert, Namibia, May.
Cheetah {Acinonyx jubatus} pawprint, Cheetah Conservation Fund, Namibia
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Suzi Eszterhas / Cheetah {Acinonyx jubatus} pawprint, Cheetah Conservation Fund, Namibia
Saami / Laplander guide shows ecotourists wolverine tracks in snow, Sweden.
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Staffan Widstrand / Saami / Laplander guide shows ecotourists wolverine tracks in snow, Sweden.
Male Leopard tracks and faeces {Panthera pardus}, early morning Malamala GR South Africa Winter
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Richard Du Toit / Male Leopard tracks and faeces {Panthera pardus}, early morning Malamala GR South Africa Winter
African lion track in wet sand {Panthera leo} Kaokoland, Namibia
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Tony Heald / African lion track in wet sand {Panthera leo} Kaokoland, Namibia
Desert Lion footprints in sand {Panthera leo} Hoarusib river, Skeleton Coast NP, Namibia
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Richard Du Toit / Desert Lion footprints in sand {Panthera leo} Hoarusib river, Skeleton Coast NP, Namibia
Lion tracks in sand Clay castles, Skeleton coast, Namibia
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Pete Oxford / Lion tracks in sand Clay castles, Skeleton coast, Namibia
Hippo spoor. Moremi reserve, Botswana, Southern Africa
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Richard Du Toit / Hippo spoor. Moremi reserve, Botswana, Southern Africa
Lion tracks (Panthera leo). Savuti-Chobe NP< Botswana, Southern Africa
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Richard Du Toit / Lion tracks (Panthera leo). Savuti-Chobe NP< Botswana, Southern Africa
Downy birch (Betula pubescens) nearly felled by a Eurasian beaver (Castor fiber). Shores of Loch Aigas, Aigas Field Centre, near Inverness, Scotland, UK. October.
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Nick Garbutt / Downy birch (Betula pubescens) nearly felled by a Eurasian beaver (Castor fiber). Shores of Loch Aigas, Aigas Field Centre, near Inverness, Scotland, UK. October.
Signs of European Beaver (Castor fiber) activity, chewed trunks of birch trees in boggy woodland. Home of a successful Beaver re-introduction project at Knapdale Forest, Argyll, Scotland, UK, Septembe...
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John Waters / Signs of European Beaver (Castor fiber) activity, chewed trunks of birch trees in boggy woodland. Home of a successful Beaver re-introduction project at Knapdale Forest, Argyll, Scotland, UK, Septembe...
Signs of the presence of European beavers (Castor fiber) including chewed tree trunks in a boggy birch woodland. Site of successful re-introduction, Knapdale Forest, Argyll, Scotland, UK, September.
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John Waters / Signs of the presence of European beavers (Castor fiber) including chewed tree trunks in a boggy birch woodland. Site of successful re-introduction, Knapdale Forest, Argyll, Scotland, UK, September.