Ernie Janes / London red bus going past Big Ben and the Houses of Parliament at Westminster, London, June 2013.

Ernie Janes / Abstract photo of Tower Bridge, with blurred motion red bus, London, June 2013. Digitally manipulated.

Jack Dykinga / Bottlenose dolphins (Tursiops truncatus) near Isla Animas, Sea of Cortez, Baja Sur, Mexico.

Bert Willaert / View from the top of the Ozarde waterfalls near Koyna Lake during the monsoon. Western Ghats, India, August 2010.

Kirkendall-Spring / Close up on mountain biker's foot pedalling, in Cascade foothills, Washington, USA. April 2013. Model released.

Chris & Monique Fallows / Common dolphin (Dephinus delphis) 'mega pod', False Bay, Cape Town, South Africa.

Denis-Huot,Denis-Huot / Lion (Panthera leo) male shaking his head after rain, Masai-Mara Game Reserve, Kenya

Ben Hall / Black headed gull (Chroicocephalus ridibundus) taking flight, artistically blurred photograph. Cheshire, UK, October. Highly honoured in the Art in Nature category of the Nature's Best Windland Smith...

Ben Hall / Black-headed gull (Chroicocephalus ridibundus) in flight, artistically blurred photograph, Cheshire, UK, February. Highly Commended in the British Wildlife Photography Awards (BWPA) competition 2013....

Dr. Axel Gebauer / Oxeye daisy (Leucanthemum vulgare) abstract, Upper Lusatia Heath and Pond Landscape, Upper Lusatia UNESCO Biosphere Reserve, Germany

Juan Carlos Munoz,Juan Carlos Munoz / Black throated diver (Gavia arctica) displaying, Norway, June.

Juan Carlos Munoz / Two juvenile Atlantic yellow legged gulls (Larus michahellis) in flight, Izmir, Turkey, October.

Juan Carlos Munoz / Male African lion (Panthera leo) at night, Kalahari Desert, Botswana.

Juan Carlos Munoz / Atlantic Puffin (Fratercula arctica) in flight, Svalbard, Norway, June.

Markus Varesvuo / Large flock of Kittiwakes (Rissa tridactyla) flying over the sea, Vardo, Norway, March.

Theo Bosboom,Theo Bosboom / Icy waterfall, Getzbach, Belgian Ardennes, February 2008

Claudio Contreras,Claudio Contreras / Black-axil Chromis (Chromis atripectoralis) blurred motion photograph. Bilang Bilangang Island, Danajon Bank, Central Visayas, Philippines, April

Cyril Ruoso / Chimpanzee (Pan troglodytes) using tools to crack nuts, Gabon

Yukihiro Fukuda / Japanese fireflies (Luciola cruciata) in flight at night, Japan endemic species, Ichikawa-Riveer, Asago Hyougo, Japan, July

David Tipling / Blue and Yellow Macaws (Ara ararauna) in flight, Tambopata Amazon, Peru

Bence Mate / European starlings (Sturnus vulgaris) taking off from waters edge after bathing, Pusztaszer, Kiskunsagi National Park, Hungary

Denis-Huot / Cheetah (Acinonyx jubatus) juvenile chasing a zebra (Equus quagga) herd, Masai-Mara Game Reserve, Kenya. Vulnerable species.

Richard Du Toit / Red billed Quelea (Quelea quelea) blurred motion flock in flight, Mpumalanga Province, South Africa, March 2012

George Sanker / Ellis River high in White Mountain National Forest. New Hampshire, October 2012.

Gavin Hellier / Opened in 2010, the Dubai Metro, MRT, in motion along Sheikh Zayed Road at dusk, Dubai, United Arab Emirates. No release available.

Gavin Hellier / In the famous souks of Marrakech, Morocco, 2011. No release available.

Gavin Hellier / Sheikh Zayed Rd, traffic and new high rise buildings along Dubai's main road, United Arab Emirates 2011. No release available.

Gavin Hellier / The restored Souq Waqif with mud rendered shops and exposed timber beams at night, Doha, Qatar, Arabian Peninsula, 2011. No release available.

Gavin Hellier / Moving train pulling into the station with people waiting, modern subway station, Berlin, Germany, 2009. No release available.

Gavin Hellier / Traffic lights and office buildings illuminated at dusk, part of new urban development, modern architecture skyscrapers in Potsdamer Platz, Berlin, Germany, 2007. As was the case in much of Berlin,...

Gavin Hellier / Khao San Road at dusk, Banglamphu, Bangkok, Thailand, 2010. No release available.

Gavin Hellier / Tuk Tuk or auto rickshaw in motion at night, Bangkok, Thailand, 2010. Model released.

Ross Hoddinott / Vehicle tracks in field of ripe Barley, farmland, late evening light, near Putford, Devon, UK. August 2012.

Jabruson / African Forest elephant (Loxodonta africana cyclotis) subadult scenting air, blurred motion at dusk, Dzanga Bai, Dzanga-Ndoki National Park, Central African Republic

Oriol Alamany / Nigel Creek with conifer forest at dusk, Banff National Park, Rocky Mountains, Alberta, Canada

Claudio Contreras / Harbour Seal (Phoca vitulina), near San Quintin, Baja California Peninsula, Mexico, December.

Claudio Contreras / Green Turtle (Chelonia mydas) hatchling on its way to the sea, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, August. Endangered species.

Oriol Alamany / African Elephant (Loxodonta africana) browsing, Kruger National Park, Transvaal, South Africa, September.

David Pattyn / Black-headed gull (Chroicocephalus ridibundus) in flight above Wadden sea, Terschelling Island, The Netherlands, June

David Pattyn / Bewick's swans (Cygnus bewickii) four in flight, Den Bosch, The Netherlands, January

Andrew Parkinson / Black-headed gull (Chroicocephalus ridibundus) adult in winter plumage, taking off at dawn from tranquil lake. (Non-ex) Derbyshire, UK. Highly commended, GDT competition 2012, Nature's studio categor...

Tui De Roy / White bellied woodstar (Chaetocercus mulsant) three hovering around red flowers, Guango private reserve, Papallacta Valley, Andean Cloud Forest, East slope, Ecuador

Philippe Clement / Great Spotted Woodpecker (Dendrocopos major) male drumming on tree trunk in forest, Belgium, July

Danny Green / Brown Bear (Ursus arctos) shaking water from fur. Finland, Europe, June.

Rob Jordan / 2020VISION / Arctic tern (Sterna paradisaea) in flight at twilight using slow shutter speed to accentuate blur and motion in low-key light, Inner Farne, Farne Islands, Northumberland, July.

Dave Watts / Tasmanian Devil (Sarcophilus harrisii) running blur, Cradle Mountain, Tasmania, Australia

Ernie Janes / Cley Beach with retreating wave patterns, after sunset, Norfolk, UK January

Ernie Janes / Indian rhinoceros (Rhinceros unicornis) charging, captive

Nick Turner / Couple with child in seat cycling in the Cotswolds at Guiting Power, Gloucestershire, UK, August 2011. Model released

Sue Flood - TD / Grey Heron (Ardea cinerea) taking a fish from water. Kiskunsagi National Park, Hungary, May.

Graham Eaton / Underwater view of a rapidly flowing mountain stream, Snowdonia NP, Gwynedd, Wales, UK, November

Graham Eaton / A split-level view through a small waterfall, Llanberis, Snowdonia NP, Gwynedd, Wales, UK, July

Graham Eaton / Underwater view of River Glaslyn rapids, Snowdonia NP, Gwynedd, Wales, UK, April

Ben Hall / 2020VISION / Stock Ghyll waterfall in autumn, with motion blur from wind, Lake District NP, Cumbria, England, UK, November

Inaki Relanzon / Trashumancia / seasonal migration, large herd of moving livestock, French Pryrenees, Spain, June 2011.

Todd Mintz / Great Horned owl, (Bubo virginianus) in flight, Regina, Saskatchewan, Canada, March

Markus Varesvuo / Dunlin (Calidris alpina) flock in flight, Finland July

Edwin Giesbers / Two Scarlet macaws (Ara macao) flying, captive

Juan Carlos Munoz / Jersey tiger moth (Euplagia quadripunctaria) large numbers resting on the trunk of Sweetgum trees (Liquidambar orientalis) some with wings open in flight, in the Petaloudes Valley, Rhodes island, Gree...

Gary K. Smith / Domestic sheep (Ovis aries) being moved from pen at livestock market after auction, Hawes Mart, Yorkshire, UK, September 2011

Sven Zacek / Waves washing up a beach under dark clouds. Hiiumaa Island, Estonia, Europe, August 2011.

Sven Zacek / Waves washing up a beach on the Baltic coast of Estonia under sunset lit clouds. Europe, August 2011.

Ole Jorgen Liodden / King Penguin (Aptenodytes patagonicus) parents calling and performing 'statue' display at colony. South Georgia Island, Southern Ocean, November.

Ole Jorgen Liodden / Black-legged Kittiwake (Rissa tridactyla) taking off from water. Spitsbergen, Svalbard, August.

Ole Jorgen Liodden / Oryx (Oryx gazella) in motion. Kalahari, Botswana, November.

Florian Möllers / African Lions (Panthera leo) in a bare cage in Beijing Zoo, China, November.

Andrew Parkinson / Whooper swan (Cygnus cygnus) adult aggressively chasing another, UK visitor between November and March. Lancashire, UK, October.

Andrew Parkinson / Jackdaws (Corvus monedula) abstract view of flock in flight, Derbyshire, UK, December.

Andrew Parkinson / Pink footed geese (Anser brachyrhynchus) huge flock take flight in dawn light, Norfolk, UK, January.

Sandesh Kadur / Termites (Isoptera) flying upwards towards a street lamp, India

Anup Shah / Leopard running - panned effect (Panthera pardus) Masai Mara National Reserve, Kenya

Bence Mate / Great reed warbler (Acrocephalus arundinaceus) taking fish from water, Hungary, June. Winner, Eric Hosking award, 2011 Wildlife Photographer of the Year competition

Bence Mate / Hoopoe (Upupa epops) in flight, Hungary. Multi flash, long exposure. Winner, Eric Hosking award, 2011 Wildlife Photographer of the Year competition.

Bence Mate / Green crowned brilliant hummingbird (Heliodoxa jacula) male taking off to feed at feeder in garden, another waiting perched on lichen covered branch, Costa Rica, January. Winner, Eric Hosking award at...

Ben Hall / Dipper (Cinclus cinclus) perched on moss-covered waterfall, Peak District NP, Derbyshire, UK, May, Highly Commended, Habitat category, British Wildlife Phototgraphy Awards (BWPA) competition 2011

Juergen Freund / Lunar tailed rock cod / Coronation grouper (Variola louti) swimming fast over coral reef, Bali, Indonesia

Simon Colmer / Wildflowers, including Poppies (Papaver) flowering, planted in central reservation / road verge, Brighton, Sussex, UK, June 2010

Alex Mustard / In-camera motion blur image of a female Hedgehog Seahorse (Hippocampus spinosissimus). Lembeh Strait, Sulawesi, Indonesia, Molluca Sea, March.

Juergen Freund / Leatherback turtle (Dermochelys coriacea) baby speeding towards the sea. Warmamedi beach, Bird's Head Peninsula, West Papua, Indonesia, July 2009

Florian Möllers / European rabbit (Oryctolagus cuniculus) near busy road, Haselhorst, Berlin, Germany, June 2008

Orsolya Haarberg / Dying Northern fulmar (Fulmarus glacialis) on the dark bottom of a deep gorge, Saefellsnes peninsula, Iceland, September 2010

Orsolya Haarberg / Coastal landscape at Dyrholaey, Iceland, April 2010

Matthias Breiter / Ruffed Grouse (Bonasa umbellus) male drumming, Kenora, Ontario, Canada

Juan Carlos Munoz / South African fur seal (Arctocephalus pusillus pusillus) swimming near surface, Seal Island. False Bay, South Africa, July 2010

Alex Mustard / A bull California sealion (Zalophus californianus) bursting through a school of baitfish in the shallows. Los Isotes, La Paz, Mexico. Sea of Cortez, East Pacific Ocean.

Edwin Giesbers / Termite (Hospitalitermes sp.) colony, with individuals moving in different directions, Sarawak, Borneo, Malaysia

Juan Carlos Munoz / Huge flock of Bramblings (Fringilla montifringilla) flying to roost in trees, Gorbeia Natural Park, Basque country, Spain, January 2011, unusual winter visitors pushed further south than usual by the...

Juan Carlos Munoz / Huge flock of Bramblings (Fringilla montifringilla) in flight, Gorbeia Natural Park, Basque country, Spain, January 2011, unusual winter visitors pushed further south than usual by the harsh winter co...

Mary McDonald / Flock of Gray-headed Gulls (Chroicocephalus cirrocephalus) flying over Lake Nakuru, Kenya, East Africa

Alex Mustard / Caribbean reef shark (Carcharhinus perezi) cruises over a coral reef. Near Grand Bahama, Bahamas. Tropical West Atlantic Ocean.

Alex Hyde / Giant millipede {Aphistogoniulus sp} running across leaf litter on rainforest floor. Andasibe-Mantadia National Park, Eastern Madagascar.

Russell Cooper / Honey bee (Apis mellifera) feeding from Lavender (Lavandula) flowers, Mayfield Lavender Farm, North Downs, Surrey, UK, July

Claudio Contreras / Guadalupe fur seal (Arctocephalus townsendi) pup swimming, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, April

Solvin Zankl / Mature female Olive ridley sea turtles (Lepidochelys olivacea) arriving on the beach of Ostional, Costa Rica, Pacific coast, during an arribada / mass nesting event, November

Chien Lee / Stream flowing through pristine lowland rainforest in Gunung Mulu National Park, Malaysia

Edwin Giesbers / Sri Lankan Leopard (Panthera pardus kotiya) soft focus running, captive.