Yukihiro Fukuda / Japanese giant salamanders (Andrias japonicus) males in a burrow at the time of spawning, Kurokawa River, Hyougo, Japan, September

Yukihiro Fukuda / Japanese giant salamander (Andrias japonicus) eating the eggs of another salamander, Japan, August

Yukihiro Fukuda / Japanese giant salamander (Andrias japonicus) moving upstream to spawn, Japan, August

Yukihiro Fukuda / Male Japanese giant salamander (Andrias japonicus) in river, Japan, August

Yukihiro Fukuda / Male Japanese giant salamander (Andrias japonicus) protecting his eggs inside his nest, Japan, October

Yukihiro Fukuda / Male Japanese giant salamander (Andrias japonicus) protecting his eggs inside his nest, Japan, September

Yukihiro Fukuda / Japanese giant salamander (Andrias japonicus) eggs, Japan, September

Yukihiro Fukuda / Japanese giant salamander (Andrias japonicus) at the entrance to its nest, raising its head to the surface, Japan, August

Yukihiro Fukuda / Japanese giant salamander (Andrias japonicus) in river, Japan, March

Yukihiro Fukuda / Japanese giant salamander (Andrias japonicus) reaching up to the surface to breathe, Japan, August

Yukihiro Fukuda / Japanese giant salamander hunting at night in a river, Japan, May

Yukihiro Fukuda / Two male Japanese giant salamanders (Andrias japonicus) fertilizing the eggs of a female, Japan, September